10 



the retractor of the right ommatophore (RRO) (Figs. 10, 2). 

 Ortiz & Ortiz (1959) missed the diminutive PR in their 

 dissections and do not show it in their illustrations. Emerging 

 above the PS, the apical vas deferens (AVD) is a large 

 muscular thick-walled conduit that in its normal position 

 reaches to the peniovaginal angle and doubtless serves both 

 as an ejaculatory duct and a physical support for the intromit- 

 tent organ. Apical to this, the conduit narrows to half the 

 calibre and is thinner walled. 



The vagina (V) is very short, but two to four times wider 

 than its length. Near its base, sparse, thin muscle strands 

 suggest a primordial vaginal retentor. Internally, the V is 

 muscular, thick-walled and longitudinally deeply plicate. 

 There is no sharp delineation between it and the broad, 

 somewhat thinner walled basal spermathecal duct (SD). This 

 latter is so large that it tends to be positioned partly between 

 P and V. Both upper SD and free oviduct (FO) are thin- 

 walled and of about the same calibre. The clavate spermath- 

 eca (S) is broadly attached to the spermoviduct apical to the 

 AVD/FO junction. The ovotestis acini are as in C. barriana. 

 No specimen was found to be gravid, but a single specimen 

 collected in October in Bonge seemed to be near it with a 

 very large albumen gland and an inflated spermoviduct. Ortiz 

 & Ortiz (1959) examined a single specimen from Fernando 

 Poo Island (Macfas Nguema Biyogo) and found the sper- 

 moviduct completely crowded with four comparatively large 

 white eggs. Seven dried eggs (MCZ no. 219224) measured in 

 the present study average 4.7 x 3.7 mm. A diminutive talon 

 is present. In contrast to C. barriana, the genital atrium is an 

 inconspicuous dimple without superficial embellishments. 



Type material. Pfeiffer (1856) described this species from 

 Cuming collection specimens, giving the shell size as 5 1 /: 

 whorls and the length-width measurements as 34 x 17 mm; 

 later (1859) he gave aperture measurements as 19 x 

 11.5 mm. The measurements of the three syntypes in BMNH 

 do not match those of Pfeiffer, but are reasonably close. The 

 largest syntype has a damaged and repaired last whorl and the 

 next largest is atypically slender; therefore the smallest 

 specimen (Figs. 25, 26; Table 2) is here selected as the 

 lectotype, the other two becoming paralectotypes. No con- 

 vincing evidence was found that other syntypes are extant. 

 Only four other specimens of the 60 examined in the present 

 study exceeded 30 mm in shell length. 



Type locality. The syntypes in the Cuming collection were 

 reported to be from ' "Grand Bassam" Africae occidentalis 

 (Verreaux)'. All other specimens examined bearing this 

 locality were sold by shell dealers, viz. Da Costa, Fulton, 

 Geret, Paetel and Preston, who may have taken their cue for 

 a locality from the original description. No museum specimen 

 has been found with a locality record from the 1400 km 

 stretch of continental Africa between Grand Bassam, Ivory 

 Coast and the cluster of reliable locality records in northwest 

 Cameroon. It is suspected that this is a case of still another 

 erroneous Cuming record. Although under the circum- 

 stances, we must accept the type locality as 'Grand Bassam', 

 it is probable that Cuming's specimens came from Cameroon, 

 or perhaps Gabon. The likelihood of an early secondarily 

 established population in Grand Bassam prior to 1856 is 

 extremely remote, for surely authentic collecting records 

 would have appeared in the meantime. Edouard Verreaux (cf 

 Crosse & Fischer, 1869) not only collected the syntypes of 

 this species, but he also collected the single known specimen 



A.R. MEAD 

 Table 2 C. shuttleworthi - Representative shells measurements. 



Greatest Aperture Last % 

 Whorls Length Width Length Width whorl LW/L % W/L 



6 



34.0 



18.7 



18.4 



10.3 



26.4 



78 



55 



'G.Bassam' 



(MCZ) 

 83441 



6 



32.3 



18.4 



18.4 



10.0 



24.5 



76 



57 



'G.Bassam' 



(BMNH) 



PLec 



6 



31.0 



16.7 



16.4 



9.0 



22.8 



73 



54 



'G.Bassam' 



(BMNH) 



PLec 



5% 



30.9 



18.3 



17.0 



10.7 



24.0 



78 



59 



'G.Bassam' 

 (BMNH) 

 Lect A . 

 shuttle- 

 worthi* 



5% 



26.9 



15.8 



14.8 



8.0 



20.7 



77 



59 



Edea 

 (MCZ) 



5'/2 



24.1 



14.8 



14.4 



8.0 



19.0 



79 



61 



Gabon 

 (MRAC) 

 5314 

 (Preston) 



5'/: 



22.9 



13.0 



12.1 



6.8 



17.3 



75 



57 



Bibundi 

 (SMNH) 



5V< 



20.2 



11.5 



11.8 



6.8 



16.1 



80 



57 



Itoki 

 (UUZM) + 



5 



19.4 



11.4 



11.2 



5.9 



15.4 



79 



59 



Bibundi 

 (SMF) 



4V2 



16.0 



10.5 



8.9 



5.3 



12.5 



78 



66 



Gabon 

 (IRSN) 

 (Vignon) 



Total specimens examined: 60. Sources: BMNH. GNM. IRSN. MCZ. MNHN. 

 MRAC. NHMW, SMF. SMNH. UMMZ. USNM, UUZM. ZMB. 



of C. barriana from Gabon, now in Bern (NHMB). This 

 raises the suggestion that Verreaux, after collecting in Gabon 

 and Cameroon, shipped his specimens from Grand Bassam, 

 which Cuming assumed was the collecting site. 



Distribution. Leonardo Fea was the first to discover this 

 species on Fernando Poo Island (= Macias Nguema Biyogo) 

 of Equatorial Guinea 3° 30' N, 8° 40' E (Germain, 1916:249. 

 Subsequently, Ortiz & Ortiz (1959:45) reported it from Basile 

 and Mongola on that island. Nine reliable localities on the 

 mainland in Cameroon define a limited coastal belt, ca 280 x 

 120 km with N'dian 4° 55' N, 8° 53' E in the north; Metet 3° 

 05' N, 11° 00' E in the east; and Itoki 2° 24' N, 9° 50' E in the 

 south. Other Cameroon localities: Albrechts Hohe 4° 38' N, 

 9° 25' E; Mukonje (= Mukonye) 4° 37' N 9° 30' E; Bibundi 4° 

 13' N, 8° 59' E; Edea 3° 48' N, 10° 08' E; Lokoundje 3° 13' N, 

 9° 55' E. A specific locality record for Gabon was not found, 

 but Vignon, through Ancey (1888:69), reports them as rare in 

 Gabon at the edge of forest streams. They probably do not 

 extend south of the Ogooue River. 



Callistoplepa tiara Preston, 1909 - A Misidentification 



Preston (1909:183, pi. vii, fig. 9) described Callistoplepa tiara 

 from 'Bitze [= Bitye], near the River Ja [Dja], Cameroons' 

 (3° 01' S, 12° 22' E). He indicated neither the collector nor 

 the number of specimens he had; however, some specimen 

 labels (BMNH, MRAC) specify that G.L. Bates was the 

 collector. Between 1908 and 1912, Preston distributed ten 

 known syntypes, each bearing the full locality information 



