26 



A.R. MEAD 



two proposed taxa. These are in the type collection in 

 Bruxelles (IRSN) and are here selected as lectotypes of 

 Dautzenberg's Achatina marteli and A.m. pallescens, respec- 

 tively (Figs. 37-40; Table 7). As he pointed out in a footnote 

 in the original descriptions, the pronounced flame pattern of 

 his figure 1 unfortunately did not reproduce well. Pilsbry's 

 copies (1905) therefore reflected this deficiency. In this 

 species, neither the lack of colour pattern nor the greater 

 degree of ventricosity is taxonomically valid for establishing a 

 trinomen. 



In the IRSN collection there are several mixed lots totalling 

 48 mostly juvenile, damaged or weathered specimens. All 

 these specimens were very carefully examined in the present 

 study and were found to be a mixture of the flamed and 

 unicolorous forms of this species and, in addition, juveniles of 

 Bequaertina pellucida and Achatina craveni. These cannot 

 reasonably be considered to have been a part of Dautzen- 

 berg's type series. Dautzenberg, however, did distribute his 

 specimens widely. Those bearing the type locality and R.P. 

 Guilleme as the collector are here selected as paralectotypes. 

 The known distribution of these flamed/unicolorous speci- 

 mens are BMNH 1/0, NMW 1/1, IRSN 8/5, MCZ 2/1, MRAC 

 7/1, NMB 2/1, MNHN 6/1, NHMW 1/0. 



Table 7 B. marteli - Representative shells measurements. 



Greatest Aperture Last % 



Whorls Length Width Length Width whorl LW/L % W/L 



6V2 68.3 34.0 42.3 20.8 55.4 81 50 Mpala 



(IRSN) 

 PLec 



6V4 67.2 38.6 39.1 22.2 54.4 81 57 Mpala 



(IRSN) 

 Lect 

 A.m. p.* 



6 63.7 33.2 37.7 19.6 51.5 81 52 Mpala 



(IRSN) 

 Lect 

 A.m.* 



6'/4 62.9 32.3 36.7 18.2 49.8 79 51 Mpala 



(MCZ) 

 97929 

 PLec 



6 60.8 34.8 38.0 19.4 51.3 84 57 Mpala 



(MNHN) 

 PLec 



5% 58.4 32.3 37.3 18.5 49.0 84 55 Mpala 



(MRAC) 

 5129 PLec 



654.032.233.817.043.78160 



54.0 32.2 33.8 17.0 43.7 81 60 Mpala 



(MNHN) 

 PLec 



5% 48.4 26.9 32.1 15.0 39.8 82 55 Mpala 



(BMNH) 

 PLec 

 1937.12. 

 30.1934 



5>/2 48.4 27.9 32.0 16.7 40.5 84 58 Mpala 



(MRAC) 

 5125 PLec 



6 48.4 27.3 31.0 15.5 39.6 82 56 Mpala 



(MRAC) 

 5131 PLec 



Total specimens examined: 69. Sources: BMNH, IRSN, MCZ, MNHN, 

 MRAC, NHMW, NMB, NMW. 



Type locality. Mpala (= Pala) 6° 45' S, 29° 31' E, region of 

 Lake Tanganyika, Zaire (cf van Burggen, 1988:9). 



Distribution. All specimens examined are from the type 

 locality. Verdcourt kindly sent the author a photograph of a 

 specimen, which earlier had been identified as Achatina 

 tavaresiana (Pain & Verdcourt, 1962; Verdcourt, 1966). This 

 specimen is clearly B. marteli and establishes this species in 

 Tanzania on the eastern shore of Lake Tanganyika in the 

 Mahari Peninsula at Nkungwe (= Kungwe) 6° 07' S, 29° 48' 

 E. It was collected in 'litter in thick scrub at head of stream, 

 altitude 4500 ft'. With the many locality records known for B. 

 pellucida, including Mpala, it is strange that the distribution 

 of the present species is, to date, so contrastingly limited. 



Remarks. This is the least fragile and the most boldly 

 sculptured species in the genus. Phylogenetically, it appears 

 to stand between B. pellucida and B. pintoi. Mixed lots of B. 

 marteli, B. pellucida, and Achatina craveni suggest that these 

 species are sympatric. The juveniles in particular are confus- 

 able. By far the largest series of this species is to be found in 

 Bruxelles (IRSN). 



Bequaertina pintoi (Bourguignat, 1889) 

 Figs. 41-51 



Serpaea pintoi 



Bourguignat, 1889:86, pi. 4. fig. 4. 

 Achatina fragilis 



Smith, 1899:591, pi. 35, figs. 3,4 (non Achatina fragilis 



Deshayes, 1864); Ancey, 1902:278. text fig. 6; Pilsbry, 



1904:63, pi. 9, figs. 25, 26 (ex Smith); Dautzenberg & 



Germain, 1914:26. 

 Achatina . . . . sp. nov?' 



Ancey, 1902:277, text fig. 4. 

 Achatina pintoi 



Pilsbry, 1904:63, pi. 41, fig. 8 (ex Bourguignat); Bequaert, 



1950:11; Verdcourt, 1966:111, 1983:219. 

 Achatina nyikaensis 



Pilsbry, 1909:113, 1919:79; Connolly, 1925:168, 1939:321; 



Germain, 1935:9; van Bruggen, 1965:81, 1988:10; van 



Bruggen & Meredith, 1984:161. 

 Callistoplepa nyikaensis 



Bequaert & Clench, 1934c:115, 116. 

 Callistoplepa thielei 



Bequaert & Clench, 1934c: 115, pi. 2, figs. 8-10, 12. 

 Callistopepla nyikaensis 



Verdcourt, 1966:111, 1983:219; Meredith, 1983a:29, fig. 



10, 1983b:247. 



Shell. Shell thin, fragile, highly variable in shape, usually 

 elongate-ovate or ovate-subsuccineiform, but may be ovate, 

 globose-ovate or slender conic-ovate. Whorls 6— 6'/2, rarely 

 slightly larger. Apex obtuse; vaguely mammillate in some 

 specimens. Spire usually inscribes a short broad based tri- 

 angle that appears to be nearly equilateral, 27-36% of shell 

 length. Less commonly, the spire is more produced, with the 

 sides of the connate triangle appearing longer than the width 

 of the base, 37-43% of shell length; such specimens may or 

 may not have a more slender last whorl. Sutures fine, distinct, 

 deeply impressed, quite regular, but may be faintly crenulate 

 in the last part of the sixth whorl. Whorls slightly, moder- 

 ately, or distinctly convex, usually expanding rapidly to form 

 a large last whorl, 84% of shell length; range for 3'/2-6 3 /4 

 whorls, 78-87% (n = 87). Aperture ovate-acuminate; charac- 



