PHYLOGENY OF ARIETELLID COPEPODS 



107 



Table 2 Characters used in the cladistic analysis for genera of the family Arietellidae. Codes to 2 refer to transformation series of 

 multi-state characters; 0: plesiomorphic state; 9: missing data. 



1. Gonopore and copulatory pore sharing common opening 



2. Right and left copulatory pores 



3. Lengths of right and left antennules of female 



4. Fusion of female antennulary segments I-III and IV 



5. Fusion between female antennulary segments XXIII and XXIV 



6. Aesthetasc present on female antennulary segment IV 



7. Aesthetasc present on female antennulary segment VI 



8. Aesthetasc present on female antennulary segment VIII 



9. Aesthetasc present on female antennulary segment X 



10. Aesthetasc present on female antennulary segment XII 



1 1 . Modification of seta into process on male antennulary segment XV 



12. Fusion of male antennulary segments XXI & XXII 



13. Seta adjacent to aesthetasc on male antennulary segments II 



14. Seta adjacent to aesthetasc on male antennulary segment III 



15. Modification of seta into process on male antennulary segment XXII 



16. Process on male antennulary compound segment XXIV-XXV 



17. Seta on first endopod segment of antenna 



18. Proximal inner seta on mid-margin of second endopod segment of antenna 



19. Vestigial element on second endopod segment of antenna 



20. 1-segmented, rudimentary mandibular endopod with 1 or 2 setae 



21. Outer terminal seta on fifth exopod segment of mandible 



22. Process on maxillulary praecoxal arthrite 



23. Inner basal enditic seta of maxillule 



24. Third seta of maxillulary endopod 



25. Inner angle seta of maxillulary endopod 



26. Distal seta on first on first praecoxal endite of maxilla 



27. Reduction of seta a on sixth endopod segment of maxilliped (length of seta 

 at most as long as the segment) 



28. Reduction of seta b on sixth endopod segment of maxilliped (length of seta 

 at most as twice as long as segment) 



29. Proximal spine on outer margin of third exopod segment of leg 1 



30. Inner coxal seta of leg 4 



31. Fusion between endopod and basis of female leg 5 



32. Inner margin of endopod of female leg 5 with proximal (seta A) and 

 distal (seta B) setae 



33. One seta (A or B) on inner margin of endopod of female leg 5 



34. Inner angle seta on distal margin of endopod of female leg 5 



35. Exopod segment of female leg 5 



36. Spine (element d) on outer distal angle of exopod of female leg 5 



37. Left endopod of male leg 5 (including incomplete fusion) 



38. Right endopod of male leg 5 



39. Seta c on third exopod segment of left male leg 5 



40. Setae e and f of left male leg 5 transformed into bifid process 



41. Third exopod segment of left male leg 5 rotated so that vestigial outer 

 margin elements now on inner surface 



42. Seta f on third exopod segment of right male leg 5 



43. Seta c on third exopod segment of right male leg 5 



44. Fusion between coxa and basis of right male leg 5 



yes/no 



O'l 



separate/fused 



1 



equal/uneuqal 



O'l 



separate/fused 



0/1 



separate/fused 



O'l 



present/absent 



( 1 



present/absent 



( 1 



present/absent 



0/1 



present/absent 



0/1 



present/absent 



1 



no/yes 



0/1 



separate/fused 



01 



present/absent 



O'l 



present/absent 



O'l 



no/yes 



O'l 



no/yes 



€■'1 



present/absent 



O'l 



present/absent 



C 1 



present/absent 



0/1 



present/absent 



1 1 



normal/reduced 



( 1 



present/absent 



( 1 



present/absent 



0/1 



present/absent 



O'l 



present/absent 



0/1 



present/absent 



0/1 



no/yes 



0/1 



no/yes 



( 1 



present/absent 



("1 



present/absent 



0/1 



separate/fused 



0.1 



A + B present/ 



t'l 



A or B absent 





present/absent 



0/1 



present/absent 



0/1 



partly defined/ 



0/1/2 



unsegmented/absent 





present/absent 



0/1 



2-segmented/ 



0/1/2 



1-segmented/absent 





1-segmented/absent 



CI 



present/absent 



0/1 



no/yes 



0/1 



no/yes 0/ 1 



well developed/minute 071 



present/absent 0/1 



separate/fused C/l 



Table 3 Character matrix for analysis using PAUP 3.0. 



Crassarietellus 



10000000000 



100100 10000100000101100001000990 



Paramisophria 



1110 10 



1 1 10 000000000000001 1 1 10001 10000 



Metacalanus 



10 110 11110 



11110 110 111110 111111112 110 10 



Arietellus 



Paraugaptilus 



Scutogerulus 



Sarsarietellus 



Pilarella 1 



Campaneria 9 9 9 



Paraugaptiloides 9 9 9 



110 111 



111111100110110111 



110111001100110111 



100111111111110111 1111111110 11112 2 10 111111 



0000010000 

 1 10 10 



10 

 9 9 9 9 

 9 9 9 9 



9 9 9 9 9 

 9 9 9 9 9 

 9 9 9 9 9 



10 10 

 10 

 10 1 



1 1 

 



1 1 

 



1 9 



10 10 



1110 

 10 9 



1 1 1 







1 1 



10 10 1 

 1 10 1 





 



10 19 9 9 9 9 

 100099999 



9 

 9 

 1 9 9 

 9 



9 9 9 9 9 9 



9 9 9 9 9 9 



9 9 9 9 9 9 



10 110 



9 1 1 



1 1 



