112 



S. OHTSUKA, G.A. BOXSHALL AND H.S.J. ROE 



Fig. 3 Crassarietellus huysi gen. et sp. nov., female. SEM micro-graphs of remnant of spermatophore attached to genital double-somite of 

 female. A, Spermatophore remnant penetrating copulatory pore, scale bar = 20 u.m; B, Spermatophore remnant, scale bar = 20 u.m. 



ethanol, BM(NH) 1993. 425. Paratype 2: £, the same collec- 

 tion date and locality as in paratype 1, only legs 2 and 3 

 dissected and mounted on glass slides, urosome mounted on 

 stub for SEM examination, prosome preserved in 70% etha- 

 nol, BM(NH) 1993. 426. 



Body length. 3.1 

 (paratypes). 



mm (holotype); 3.88, 3.85 mm 



Description. Female. Body (Fig. 1A,B) oval in dorsal 

 view. Cephalosome and first pedigerous somite separate; 

 fourth and fifth pedigerous somites completely fused; poste- 

 rior corner of prosome produced posteriorly into rounded 

 lobe directed backwards, reaching half length of genital 

 double-somite. Urosome (Fig. 1C) 4-segmented, one-third as 

 long as prosome; genital double-somite (Figs 1D,E,2A,B) 

 wider than long; pair of medial gonopores (Fig. 2C,D) 

 located ventro-laterally near mid-level of double-somite; 

 paired copulatory pores posterior to gonopores, each con- 

 cealed beneath ventrolateral projection; remnants of diver- 

 gent fertilization tubes of spermatophore (Fig. 3) still 

 attached to genital double-somite of both paratypes, each 

 connecting through posteroventral groove with copulatory 

 pore beneath projection; copulatory duct swollen in ventro- 

 lateral projection, almost horizontal, extending to large semi- 

 nal receptacle; 1 medial and 2 pairs of lateral shallow 

 chitinized pits anteriorly; anal somite small, anal operculum 

 not developed; caudal ramus (Fig. 1C) longer than wide, 

 fringed with long setules along inner margin, with vestigial 

 seta I and developed setae II to VI, seta VII originating 

 dorsally near base of seta VI; inner margin near anus with 



patch of minute spinules. Integument of body and append- 

 ages pitted. 



Antennules (Fig. 4A-C) equal in length, distinctly 

 22-segmented, reaching to posterior end of second pediger- 

 ous somite; distal 2 segments incompletely fused; fusion 

 pattern and armature as follows: I — III— 7 + 2 aesthetascs, 

 IV-2 + aesthetasc, V-2 + aesthetasc, VI-2 + aesthetasc, 

 VII-2 + aesthetasc, VIII-2 + aesthetasc, IX-2 + aesthetasc, 

 X-2 + aesthetasc, XI-2 + aesthetasc, XII-2 + aesthetasc, 

 XIII-2 + aesthetasc, XIV-2 + aesthetasc, XV-2 + aes- 

 thetasc, XVI-2 + aesthetasc, XVII-2 + aesthetasc, XVIII-2 

 + aesthetasc, XIX-2 + aesthetasc, XX-2 + aesthetasc, 

 XXI-2 + aesthetasc, XXII-1, XXIII-1, XXIV-XXVIII-12 

 + 2 aesthetascs. Segments I to XIII fringed with long setules 

 along posterior margin; segments IV to VIII each furnished 

 with transverse row of minute setules near posterior corner. 

 Sutures between segments I to III weakly visible. 



Antenna (Fig. 1F,G): coxa unarmed; basis with spinulose 

 seta at inner angle; endopod 2-segmented, first segment with 

 minute seta at three quarters length, covered with minute 

 spinules distally, second with 3 setae of unequal lengths 

 medially and 5 setae distally and sparsely covered by spinules; 

 exopod indistinctly 10-segmented, second to fourth segments 

 almost fused; armature as follows: 0,0,0,1,1,1,1,1,0,3; ninth 

 segment sparsely ornamented with minute spinules. 



Mandible (Fig. 4D): gnathobase heavily chitinized, ventro- 

 medial margin with dense fringe of long setules; cutting edge 

 with 3 acute teeth, dorsalmost of which bifid at tip; 2 patches 

 of dagger-like spinules present dorsally; tuft of long setules 

 present medially on knob; basis of palp with patches of 



