120 



S. OHTSUKA, G.A. BOXSHALL AND H.S.J. ROE 



The male of the new genus is similar to that of Crassarietel- 

 lus. However, the left antennule, the antennary exopod, the 

 maxillulary praecoxal arthrite, and the fifth and sixth endo- 

 pod segments of maxilliped are different: (1) left antennule 

 reaching almost to end of urosome in Campaneria, but, 

 possibly, at most to end of prosome in Crassarietellus; (2) 

 antennulary segments II to IV partly fused in Campaneria, 

 but almost completely so in Crassarietellus; (3) antennulary 

 segments II and III each bearing single seta and aesthetasc in 

 Campaneria, but 2 setae and aesthetasc in Crassarietellus; (4) 

 antennulary segments XXI and XXII completely separate in 

 Campaneria, but almost fully fused in Crassarietellus; (5) seta 

 on antennulary segment XV modified into process in Cras- 

 sarietellus, but not in Campaneria; (6) antennary exopod 

 indistinctly 8-segmented in Campaneria but 10-segmented in 

 Crassarietellus; (7) spines on maxillulary praecoxal arthrite 

 finely serrate in Campaneria, but strongly serrate in Crassari- 

 etellus; (8) innermost seta on the fifth endopod segment of 

 maxilliped relatively short in Campaneria, but long in Cras- 

 sarietellus; (9) seta a on the sixth endopod segment of 

 maxilliped relatively reduced in Campaneria, but not in 

 Crassarietellus . 



The leg 5 of Campaneria is also similar to that of Crassari- 

 etellus sp., particularly in having a 2-segmented left endopod, 

 but can be distinguished by the presence of the right endopod 

 and by the 2-segmented left exopod. 



Etymology. The new genus Campaneria is named in 

 honour of the late Dr. A. Campaner who was the first to be 

 interested in the phylogenetic relationships between arietellid 

 genera (gender feminine). The specific name latipes (Latin 

 latus meaning broad; Latin pes meaning leg) refers to the 

 broad compound exopod segments of the right leg 5 of the 

 male. 



Ecological note. Campaneria was collected by a trawl 

 from the near-bottom samples taken at depths of 1234-1260 

 m off northeastern New Zealand (Bradford, 1969). Since the 

 genus has never been captured in plankton hauls, it is most 

 likely hyperbenthic. 



Campaneria latipes gen. et sp. nov. (Figs 9-10) 



Material examined, cf, New Zealand Oceanographic 

 Institute Reg. No. 121, labelled as Scutogerulus pelophilus 



(cD. 



Body length. 3.9 mm (after Bradford, 1969). 



Description. Anal somite (Fig. 9A) small, almost tele- 

 scoped into preceding somite; caudal rami (Fig. 9A) sym- 

 metrical, seta I vestigial, setae II-VI developed, seta VII 

 minute. 



Left antennule (Fig. 9B-F): segment I damaged, but with 3 

 setae and aesthetasc (only this segment still remained on the 

 body); segments II and III fused with suture visible ant- 

 eriorly; segments III and IV, and XXIV-XXV and 

 XXVI-XXVIII incompletely fused. Fusion pattern and arma- 

 ture elements as follows: II-IV-4 + 3 aesthetascs, V-2 + 

 aesthetasc, VI-2 + aesthetasc, VII-2 + aesthetasc, VIII-2 + 

 aesthetasc, IX-2 + aesthetasc, X-l + aesthetasc + process, 

 XI-2 + aesthetasc, XII— 1 + aesthetasc + process, XIII— 1 + 

 aesthetasc + process, XIV-1 + aesthetasc + process, XV-2 

 + aesthetasc, XVI-2 + aesthetasc, XVII-2 + aesthetasc, 

 XVIII-2 + aesthetasc, XIX-1 + aesthetasc + 2 processes, 

 XX-1 + aesthetasc + process, XXI-aesthetasc + 2 processes, 



XXII-XXIII-1 + process (XXII-process, XXIII-1), 

 XXIV-XXVIII-12 + 2 aesthetascs. 



Antenna: inner basal seta present; endopod (Fig. 9G) 

 2-segmented, first segment with short inner seta, second 

 segment with 3 inner setae of unequal lengths subterminally 

 and 5 setae terminally; exopod (Fig. 10A) indistinctly 

 8-segmented, second segment elongate, setal formula 

 0,1,1,1,1,1,0,3. 



Mandibular palp (Fig. 10E): endopod rudimentary, 

 1-segmented, carrying 2 setae of unequal lengths; first exo- 

 pod segment bearing non-reduced seta, fifth segment with 1 

 long and 1 shorter seta. 



Maxillule: praecoxal arthrite (Fig. 10B) bearing 5 spines 

 and 1 process, 3 of which serrate medially, with row of long 

 setules and patch of minute spinules proximally; coxal endite 

 (Fig. 10C) with long spinulose seta terminally; coxal epi- 

 podite with 6 setae; minute endite seta present on basis (Fig. 

 10D), endopod bulbous, 1-segmented, with 2 spinulose setae 

 of unequal lengths. 



Maxilla: first praecoxal endite with 2 spinulose setae and 

 vestigial element, second endite with 2 bipinnate setae (Fig. 

 10F); basal spine (Fig. 10G) with 3 rows of spinules of 

 different sizes proximally. 



Maxilliped: fourth and fifth endopod segments (Fig. 10H) 

 each having non-reduced, spinulose innermost seta, but seta 

 on fourth segment much longer than on fifth; sixth endopod 

 segment (Fig. 101) with medium-length seta b and vestigial 

 seta a. 



Leg 1 with 2 outer spines on third exopod segment. Leg 4 

 having outer basal seta, but lacking inner coxal seta. 



Leg 5 (Fig. 10J): coxae and intercoxal sclerite almost fused, 

 but suture visible on posterior surface; basis separate from 

 coxa. Right leg: basal seta missing; endopod 1-segmented, 

 with tuft of short setules terminally; exopod indistinctly 

 3-segmented, first triangular, carrying spine at outer angle, 

 distal 2 segments almost fused, but suture visible on both 

 surfaces, expanded medially, having outer seta proximally, 

 round inner process with 3 minute prominences at tip medi- 

 ally, and 2 setae and 1 prominence along outer terminal 

 margin. Left leg: basal seta missing; endopod indistinctly 

 2-segmented, unarmed; exopod 2-segmented, first segment 

 triangular, bearing spine on outer corner, second segment 

 expanded inwards, curved outwards at about three quarters 

 length, with fine medial seta and 3 terminal setae of unequal 

 lengths. 



Remarks. In her original description Bradford (1969) over- 

 looked the antennary basal seta, the inner seta on the first 

 antennary endopod segment, 3 short setae on the distal 2 

 endopod segments of the maxilliped, the outer basal seta of 

 leg 4, and the fine midlength seta on the second exopod 

 segment of left leg 5. 



Genus Paraugaptiloid.es gen. nov. 



Diagnosis. Only male known. Body similar to that of 

 Paraugaptilus; cephalosome separate from first pedigerous 

 somite; prosome rounded anteriorly and produced posteri- 

 orly, with small dorsolateral prominence and bluntly pro- 

 duced lateral lobe on each side; lateral flap of cephalosome 

 developed to cover bases of mouthparts. Caudal rami sym- 

 metrical with setae II and III normally developed. 



Male left antennule 19-segmented, fringed with setules 

 along posterior margin of first segment only; segments I and 



