62 



S. KARAYTUG AND G.A. BOXSHALL 



Coxa Basis Exopod 



Endopod 



Legl 



0-1 



1-1 



I-1;I-1;III,1,4 



0-1 ;0-l; 1,1,4 



Leg 2 



0-1 



1-0 



I-1;I-1;III,I,5 



0-l;0-2;l,I,4 



Leg 3 



0-1 



1-0 



I 1;I-1;III,I,5 



0-1 ;0-2; 1,1,4 



Leg 4 



0-1 



1-0 



I-1;I-1;III,1,4 



0-1 ;0-2; 1,11,2 



Leg 6 represented by 1 plumose and 1 naked seta at 

 posterolateral angle of second urosomite (Fig. 6E) 



Male Copepodid V 



Mean body length: mean ± standard deviation = 695 ± 44.83 urn 

 (range 634 to 743 um, n=7), mean body width 216 ± 8.1 urn 

 (range 200 to 221 .73 um). Differing from male Copepodid IV as 

 follows: body 9-segmented: first to fourth urosomites 

 ornamented along posterior margins (Fig. 14B). Anal somite 

 densely furnished with spinules along distal margin and 

 extending either side of anal operculum. 



Appendages as in fourth copepodid except as follows: 

 antennule (Fig. 15B) 7-segmented; proximal 3 segments much 

 enlarged, swollen (Figs 16C) first segment with one modified 

 spiniform seta (stippled on Fig. 15B); segment 3 with 6 similarly 

 modified elements and segment 4 with 1 such element. Setal 

 formula; 19+1 spine, 5, 5 + 6 spines, 2 + 1 aesthetasc + 1 spine, 2, 

 3,7+1 aesthetasc. 



Leg 6 (Fig. 14B) represented by 3 setae, not visible in dorsal 

 view. 



Adult female 



Body length: mean ± standard deviation = 935 ± 61.07 um (range 

 833 to 1013 um, n=10), mean body width 307 ± 14.9 um (range 

 291 to 330 um, n=10). Body (Fig. 17A) comprising 4-segmented 

 prosome and 5-segmented urosome. Prosome with 

 cephalothorax and 3 free pedigerous somites decreasing in width 

 from anterior to posterior. Cephalothorax narrowing anteriorly; 

 widest aproximately in middle. Third and fourth pedigerous 

 somites with lateral groups of bristles at posterolateral angles. 

 Urosome (Figs 18A,B) consisting of 5th pedigerous somite, 

 genital double-somite and 3 free abdominal somites. Genital 

 double-somite about as long as broad. Genital double-somite 

 and first 2 free abdominal somites with surface ornamentation 

 dorsally and ventrally : fifth pedigerous somite with 

 ornamentation along posterior margin. Anal somite with 

 spinular row ventrally extending round to anal operculum (Fig. 

 19A). Anal operculum smooth; row of spinules present in anal 

 cleft either side of midline (Fig. 19D). 



Caudal rami about 3.5 times longer than broad with 

 ornamentation comprising rows of pits or cuticular depressions 

 on ventral surface (Fig. 19B). Generally held wide apart, slightly 

 divergent. Caudal rami with 6 setae; seta I missing; outer lateral 

 seta (III) with spinular row at base extending dorsally and 

 ventrally; small seta (II) on dorsolateral surface with spinular 

 row behind it and extending ventrally. 



Antennule 8-segmented (Fig. 17B), quite short. First segment 

 with spinular row ventrally. Segment 2 with partial suture line. 

 Fourth segment longest. Segment 5 distinctive with short 

 aesthetasc (Fig. 16B); apical segment with aesthetasc fused to 

 adjacent seta at base (Fig. 16E). Setal formula: 8, 12, 6, 5, 2 + 1 

 aesthetasc, 2, 3, 7 + 1 aesthetasc. One element on seventh 

 segment possibly a setiform aesthetasc (see discussion below). 



Antenna 4-segmented (Fig. 8F), comprising coxobasis and 

 3-segmented endopod. Coxobasis with complex ornamentation 



as figured and armed with 2 inner setae, and 1 outer spinulose 

 seta representing exopod. First endopodal segment with inner 

 distal seta and midsurface spinular row. Second endopodal 

 segment with 9 setae, of which 5 on inner margin and 4 arranged 

 along inner part of distal margin; segment ornamented with 

 spinules along outer margin. Third endopodal segment armed 

 with 7 setae around apex; segment ornamented with spinular 

 row along outer margin. 



Labrum with complex ornamentation on ventral surface (Fig. 

 20A). Anterior part broader than posterior part. Posterior 

 margin forming strong teeth. 



Mandible (Fig. 20B) consisting of well developed coxal 

 gnathobase and reduced palp. Gnathobasic blades mostly 

 simple, dorsal seta with spinules along inner rim. Palp 

 represented by 3 setae, one long and spinulose, the other 

 plumose. Third seta short and naked. Central surface of coxa 

 with 3 spinular rows and another spinular row on margin at base 

 of gnathobase. 



Maxillule (Fig. 20C) consisting of powerful praecoxa and 

 reduced 2-segmented palp. Praecoxal arthrite armed with 7 setae 

 articulating at base and 4 spines fused to segment; proximalmost 

 seta spinulose, spines naked. Proximal segment of palp derived 

 from coxa and basis, bearing 2 naked and 1 spinulose inner 

 margin setae, plus outer seta representing exopod. Distal 

 segment of palp, representing endopod, armed with 2 setae with 

 spinules along margin and 1 naked seta. 



Maxilla 5-segmented (Fig. 20D), comprising praecoxa, coxa, 

 basis and 2-segmented endopod. Praecoxa with spinular row on 

 outer margin. Praecoxal endite with 2 setae, 1 of which 

 spinulose. Coxa with proximal endite represented by single seta, 

 distal endite with well developed process carrying strong 

 spinulose seta and naked seta apically. Basis drawn out into 

 powerful spinulate claw and armed with strong accessory claw 

 with spinular row along convex margin and naked seta. First 

 endopodal segment carrying 2 spinulose setae, second carrying 3 

 setae. 



Maxilliped 4-segmented (Fig. 20E), comprising syncoxa, 

 basis and 2-segmented endopod. Syncoxa armed with 3 inner 

 margin setae representing endites; ornamented with spinular 

 row near middle of inner margin and 2 spinules on outer margin. 

 Basis armed with 2 inner setae, 1 of which with spinules; 

 ornamented with 2 transverse rows of spinules near outer distal 

 angle and another spinular row midway along outer margin. 

 First endopodal segment bearing claw-like seta with 5 spinules at 

 midlength. Second endopodal segment with 3 setae, 2 of which 

 bearing spinules midway; outermost naked. 



Legs 1 to 4 each with complex ornamentation on anterior 

 and posterior surfaces of coxa as figured; also ornamented on 

 intercoxal sclerite. Legs 1 to 3 with spinular rows on posterior 

 surface of exopodal segments 1 and 2, and endopodal segment 

 2. 



Leg 1 (Fig. 10F) with 3-segmented protopod. Praecoxa 

 represented by triangular sclerite at outer proximal angle. Coxa 

 with inner plumose seta. Basis with outer angle seta and setiform 

 spine on inner margin; bearing spinular row on posterior 

 surface. Both seta and spine with spinular rows at bases. 



Legs 2 (Fig. 1 IF) and 3 (Fig. 12E) with 3-segmented 

 protopods. Coxa with inner setiform spine. Outer seta on basis 

 with spinular row at base. First segment of exopod with spinular 

 row on anterior surface. 



Leg 4 (Fig. 13D) with 3-segmented protopod. Praecoxa 

 represented by thin, hooped sclerite. Coxa with inner setiform 

 spine. Basis with outer seta bearing spinular row at base; 

 spinular row on inner margin anteriorly. Segments 1 and 2 of 



