104 



P.D. JENKINS AND A.L. SMITH 



Fig. 1 Dorsal view of cranium from left to right of C. attenuata BM(NH) 191 1.9.8.26, C. hilliana BM(NH) 1994. 113 and C. fuliginosa 

 BM(NH)1933.4.1.183. 



condyle width to height 91.2-113.3), the postero-internal 

 ramal fossa has a broad base and is approximately as broad as 

 deep; the mental foramen is positioned below the anterior 

 part of the first lower molar (M,). 



The dentition is illustrated in Figs 2-6. The first upper 

 incisor (I 1 ) is robust, slightly proodont with well developed 

 posterolingual and posterobuccal cingula; the upper unicus- 

 pids are overlapping and crowded; the first upper unicuspid 

 (Un 1 ) is large and broad in comparison with the other 

 unicuspids, its breadth is equal to or greater than the distance 

 between the two first unicuspids and this tooth is more than 

 two thirds the height of I 1 and P 4 ; the second and third 

 unicuspids (Un 2 and Un 3 ) are subequal in size; the upper 

 premolar (P 4 ) has a moderately small parastyle and robust 

 metacone; the first and second upper molars (M 1 and M 2 ) 

 show no significant distinguishing features; the third upper 

 molar (M 3 ) is short and slender with a slightly compressed 

 lingual basin. The first lower incisor (I : ) is robust, long, deep 

 and curved, and the anterolingual ridge extends for circa 

 three quarters of the length of the tooth, diverging from the 

 ventral border, the posterior border of I x lies below the 

 middle of the lower premolar (P 4 ); two thirds of the second 

 lower incisor (I 2 ) are in contact with I, and one third of the 

 tooth is overlapped by P 4 ; the postentoconid ledge is very 

 narrow in the first lower molar (M,) and yet more reduced in 

 the second lower molar (M 2 ); the talonid of the third lower 



molar (M 3 ) is reduced to a single cusp. 



Etymology 



This species is named in honour of John Edwards Hill, who 

 taught one of the authors (PDJ) the basics of mammalogy and 

 who also provided invaluable help in the identification of 

 some of the skull fragments of bats found during the survey. 



Comparison with other species 



Five species of Crocidura have been recorded from Thailand 

 (Lekagul & McNeely, 1977, Davison, 1984): C. fuliginosa 

 (including C. dracula Thomas, 1912 listed as a separate 

 species by Lekagul and McNeely), C. attenuata, C. pullata 

 vorax Allen, 1923 (listed as C. russula vorax), C. horsfieldii 

 indochinensis Robinson & Kloss, 1922 and C. monticola 

 Peters, 1870. Crocidura hilliana is separated from most 

 specimens of C. fuliginosa dracula by its smaller size (see 

 Table 1), while it is considerably larger than C. p. vorax 

 (condylobasal length <17.5), C. horsfieldii (condyloincisive 

 length <17.9, data taken from Heaney & Timm (1983) for 

 specimens from Vietnam) or C. monticola (condylobasal 

 length <17.4). 



Crocidura hilliana falls at the middle to upper part of the 

 cranial size range of C. attenuata and the lower part of the size 



