THE GERMAN CARP IN THE UNITED STATES. 583 



It seems probable, however, that the cha)-ac,tor of the scales should 

 be placed with the other four categories of characters given above 

 as being another modification brought about by artificial breeding and 

 selection and not as a condition due simpl}' to conditions of domesti- 

 cation, as is sometimes supposed. All of these characters are probably 

 heritable, although some of them, such as rate of growth and time of 

 sexual maturity, may undoubtedly be readil}' influenced b}^ external 

 conditions in the individuals of a single generation. Furthermore, 

 there apparent!}- can be all combinations of these characters, and the 

 so-called different varieties and races are the fish possessing the vari- 

 ous combinations. In general, it may be said that the most highly 

 specialized carp are those which are destitute of scales, which grow 

 quickly, are high in proportion to their length, and tend to have a hump 

 back of the head, and which become sexually mature at an early age. 



These various forms of carp probabl}^ difi'er in no essential way, 

 except that they are not so well differentiated and established, from 

 what are spoken of as "breeds" by stock breeders. There would 

 appear to be no valid reason for calling those vfith the different char- 

 acter of scales "varieties," and to class those which are differentiated 

 as to form as "races." It is merel}' that the most obvious characters 

 are those which have become most permanentl}^ established by selec- 

 tion, namely, character ol scales first and form second. Walter 

 claims that ability for quick growth has also been fixed in certain 

 stocks. Thus a fish of good quick-growing stock may later make a 

 good growth even if poorly nourished during its first or second year, 

 whereas a fish of poor stock under similar conditions would be perma- 

 nently stunted. The hardiness, or ability to resist climatic condi- 

 tions, he says has not yet been made permanent in an}^ stock, though 

 it is claimed that scale carp possess the ability to a greater degree than 

 the others. The adaptability to climatic conditions probably becomes 

 reduced rather than increased as the other characters are developed. 



All intermediate stages are found in the sets of characters men- 

 tioned. For example, fish may be entirely covered with scales, but 

 the scales are larger and fcAver in number than on the regular scale 

 carp, and, similarly, one finds all gradations between the leather and 

 the mirror carp. The same thing is true of the form of the body. 

 This is especial!}' the case with the fish in our waters, where all kinds 

 have become established and have interbred until there is a complete 

 series in the gradation of characters in almost any lot of fish taken, 

 and a division of them into varieties must be an arbitrary one. As a 

 matter of convenience in my work, those fish whicli had larger and 

 fewer scales than typical scale carp I called mirror carp. Some 

 authors state that the leather carp should l)e entirely destitute of scales; 

 others that it may have a row of scales along the back and a row on 

 each side. In no case in the Great Lakes did I see a carp entirely 



