P.R. PUGH 

nl 
Fig.3 A. upper and B. lower views of young nectophore. Scale bar 5 mm. 
[ 

Fig.4  Erenna richardi. A. first and B., C. second types of bract. D. detail of distal end of a bract. A-C. scale 2 mm; D. scale 0.2 mm. 
were situated in the distal half of the second type of bract, but in the 
first type they were positioned at about 13-14 mm from the proximal 
end of the bract, whatever the length of the latter. Thus in the longest 
bracts they were situated at about one quarter to one third its length; 
whereas in smaller ones they could be positioned in the distal half of 
the bract. Longer bracts of the first type had a second pair of lateral 
cusps close to the distal end. For both types, the bracteal canal 
originated, proximally, on the dorsal surface of the bract. For most of 
its course it remained in close contact with the ventral wall of the 
bract and there were striated bands of cells on each side of it. Close 
to the distal end of the bract it penetrated through the mesogloea to 
terminate below a small cup-shaped clump of large epidermal cells 
sunk into the dorsal surface at the tip of the bract (Fig. 4D). Some of 
these cells in this clump probably produced bioluminescence; others 
were nematocysts measuring c. 68 x 24 um. No patches of such 
epidermal cells were noted elsewhere on the bract. 
GASTROZOOID. (Fig. 5 A, B). Large gastrozooids contained dark 
brown (black in life) pigment. The proboscis region often was 
widely open and folded back on itself exposing a mass of villi. 
Enormous basigaster with two large lateral lobes, and also ex- 
panded, to a lesser extent, on the side opposite to where the tentacle 
was attached. No obvious pedicle. 
TENTACLE AND TENTILLUM. (Figs 6, 7). Bedot’s (1904) original 
description and illustrations of the tentacle and tentilla were detailed 
and accurate, and need little elaboration. The annulated tentacle had 
a muscular lamella running down one side, with the tentilla attached 
on the opposite side, at the internodes. 
Each tentillum consisted of a pedicle (Fig. 7p), a cnidoband (Fig. 
7c) and a terminal process (Fig. 7tp). The largest tentilla (Fig. 6) 
have a pedicle of up to 4-5 mm; longer than Bedot described. 
However, no doubt its length can be varied in life, and photographs 
of the specimen before preservation showed the pedicle to be highly 
contracted. The cnidoband measured up to c. 15 mm long and was 
laterally compressed. It consisted of the cnidoband proper, where 
the nematocysts are attached, and what Totton (1965) called, in the 
case of Pyrostephos vanhoeffeni Moser, 1925, the saccus (Fig. 73). 
