110 
to these six species, one, Stegocephaloides calypsonis sp.nov., is 
formally described for the first time, although it was first recognized 
by Anton Galan in his unpublished Ph.D. thesis (1984). 
As a ‘spin-off’ effect of describing some of the above mentioned 
species, six further new species are described: Andaniexis americana, 
A.gloriosa and A. pelagica, Andaniopsis africana, Stegocephaloides 
boxshalli and S. ledoyeri. All these species have previously been 
assigned to one of the six redescribed species. 
The phylogenetic relationships and generic status of the species 
described herein are not discussed in the present paper, which is one 
of a series (together with Berge & Vader 1997 a—d) that will 
collectively revise all the genera and species in the family, and 
eventually lead to a phylogenetic analysis of the Stegocephalidae 
Berge & Vader, in press). 
MATERIAL AND METHODS 
The present study is based primarily on material from the collections 
of the Natural History Museum in London, UK (BMNH). Addi- 
tional material has been borrowed from the American Museum of 
Natural History, New York (AMNH) and the South African Mu- 
seum, Cape Town (SAM). 
All dissected appendages were mounted in polyvinyl-lactophenol 
and stained with rose-bengal. Figures of these appendages were 
made using a Leica compound microscope, while the habitus- 
drawings were made using a Leica dissecting microscope. Mature 
and immature females were distinguished from males by the pres- 
ence of oostegites. 
Scales on figures are all 0.1 mm. Classification of setae and setae- 
groups follow the scheme of Berge (2001b). 
Symbols: Al—2: Antenna 1—2; EP3: Epimeral plate 3; L: Labium; 
LBR: Labrum; LMND: Left mandible; MX1: Maxilla 1; MX2: 
Maxilla 2; MXP: Maxilliped; P1—7: Pereopods 1-7; PLP: palp; 
RMND: Right mandible; T: Telson; U1—3: Uropod 1-3. 
SYSTEMATICS 
Family STEGOCEPHALIDAE Dana, 1855 
Type genus: STEGOCEPHALUS Kroyer, 1842 
Andaniexis Stebbing, 1906 
Andania Boeck, 1871: 128. Homonym, Lepidoptera 
Andaniexis Stebbing, 1906: 94 
Andaniexis australis K.H.Barnard, 1932 
(Figs 1-3) 
Andaniexis australis K.H.Barnard, 1932: 76 
Non A. australis Barnard 1964 (= A. pelagica sp.nov.) 
Non A. australis Barnard 1962 (= A. americana sp.nov.) 
Non A. australis Ledoyer 1986 (= A. gloriosa sp.nov.) 
MATERIAL EXAMINED. Syntypes 4 females, BMNH 
1936.11.2.588-591, 33-34°S 9-16°E, 1000m. 
DISTRIBUTION. Only the type material is known. 
DESCRIPTION 
Based on type material. Females 4—Smm. 
Rostrum very small. 
J. BERGE, W. VADER AND A. GALAN 




Fig. 1 Andaniexis australis, female cotype. 
Antennae short. Antenna | longer than antenna 2; flagellum 6— 
articulate; accessory flagellum article 2 absent. Antenna 2 peduncle 
(articles 3—5) shorter than flagellum; article 3 short, about as long as 
broad; article 4 shorter than article 5. 
Epistome produced laterally; rectangular with a long ridge on 
each side. 
Epistomal plate (medial keel) produced; small elongate medial 
ridge exceeding along the entire epistome. 
Mandible incisor transverse; incisor smooth; left lacinia mobilis 
present, reduced, distally straight, not conical. 
Maxilla 1 palp 2-articulate, oval, apex reaching beyond outer 
plate; outer plate distally rounded, ST in two parallel rows, first 
marginal and second submarginal, ST first row with 6 setae (ST1-5, 
ST7), ST 1 ordinary (similar to ST 2-4), gap between STS and ST7 
present, ST A-C present and part of second row; inner plate with a 
weakly developed shoulder; setae pappose. 
Maxilla 2 ordinary; outer plate setae without distal hooks or cleft; 
inner plate setae row A covering the entire margin, clearly separated 
from row B, row A setae pappose; 3-4 first setae pectinate distally; 
row B setae proximally pappose, distally with cusps present; row C 
present; row D present, 1—3 long setae distally, setae slender. 
Maxilliped palp 4-articulate, article 2 distally unproduced, dacty- 
lus distally simple (pointed); inner plate with 1 nodular seta; medial 
setae-row present, not reduced, transverse, setae pectinate; distal 
setae-row present, setae simple; inner setae-row present, but re- 
duced to one or two setae, conspicuously large and strong; outer 
plate outer setae-row present, setae submarginal, attached normally, 
long robust, straight; inner setae-row present, well developed, setae 
long robust, slender, appressed to outer setae-row; distal setae-group 
absent. 
