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Fig. 1 A. Holotype of N. tuberigum, MZB 9356, 48.5 mm SL, gravid 
female. B. Allotype of N. tuberigum, MZB 10565, 43.0 mm SL, mature 
male. Scale bar in mm. 

Station, caught in a muddy forest stream tributary to Sungai Lembang; 
R.K. Hadiaty and A. Mun’im; 4 September 1997. 
Non-type materials: MZB 9351, 10 ex., (38.0-53.0 mm SL); data 
as for holotype. MZB 9362, 28 ex., (33.2-45.9 mm SL); location as 
for holotype; R.K. Hadiaty and A. Mun’im; 31 August 1997. MZB 
9363, 27 ex., (34.7-52.2 mm SL); location as for holotype; R.K. 
Hadiaty and A. Mun’im; | September 1997. MZB 9364, 11 ex., 
(33.2-52.3 mm SL); Indonesia, Sumatra, Aceh Selatan, Desa Pucuk 
Lembang, Alur Betung, a tributary of Sungai Lembang; R.K. Hadiaty 
and A. Mun’im; 2 September 1997. MZB 10566, 2 ex., (47.2-48.2 
mm SL); unnamed tributary of S. Alas, behind the camp at Ketambe 
Research Station, Aceh, Sumatra, Indonesia; R.K. Hadiaty & A. 
Mun’im; 21 June 1998. MZB 10567, 3 ex., (38.6—52.9 mm SL); 
unnamed tributary of S. Alas, Ketambe Research Station, Aceh, 
R.K. HADIATY AND D.J. SIEBERT 
Sumatra, Indonesia; R.K. Hadiaty & A. Mun’im; 20 June 1998. 
MZB 10568, 5 ex., (41.2-52.8 mm SL); unnamed tributary of S. 
Alas, in front of the National Park camp ground, Ketambe, Aceh, 
Sumatra, Indonesia; R.K. Hadiaty & A. Mun’im; 23 June 1998. 
MZB 10569, 2 ex., (39.8-44.9 mm SL); mouth of S. Sukarimbun, 
Ketambe, Aceh, Sumatra, Indonesia; R.K. Hadiaty & A. Mun’im; 
20 June 1998. MZB 10570, 15 ex., (28.2-59.6 mm SL), unnamed 
tributary of S. Alas, behind the camp of the Ketambe Research 
Station, Sumatra, Indonesia; R.K. Hadiaty & A. Mun’im; 21 June 
1998. 
DIAGNOSIS. Nemacheilus tuberigum is easily distinguished from 
all other Nemacheilus by the following combination of characters: a 
row of comparatively large tubercles present on enlarged, elongate 
scales in the scale rows immediately above and below the lateral line 
scale row on the anterior half of the caudal peduncle, (comparatively 
large tubercles in a similar position present also in N. selangoricus 
and WN. spiniferus, but on scales with peculiar posterior elongate 
processes); and colour pattern consisting of 11-15 dorsal saddles, a 
series of 8—13 lateral blotches centered along the lateral line and 
without a dark spot at anterior base of dorsal fin (present in N. 
selangoricus and N. spiniferus). 
DESCRIPTION. General appearance and physiognomy are shown 
in Figure 1; morphometric and meristic data for the holotype and 
paratypes (range, as the minimum and maximum observation, mean 
and standard deviation) are given in Table 1. Nemacheilus tuberigum 
presents a shape and colour pattern general for many members of the 
genus. 
Dorsal head length 5—6 times in SL; eye moderate, shorter than 
snout, 34 times in dorsal head length; suborbital flaplet present in 
males at anteroventral corner of eye, anterior nares a short tube (Fig. 
2); small tubercles scattered over dorsal and lateral exposures of the 
head. Cephalic lateral line pores: supraorbital canal with 5 pores; post- 
temporal canal with 3 pores, infraorbital canal with 10 pores, supra- 
occipital canal with 3 pores; operculomandibular canal with 9 pores. 
Mouth crescent-shaped (Fig. 3), with three pairs of barbels: 
anterior rostral barbel shortest, reaching to about middle of eye; 
posterior rostral barbel longest, reaching to half way between hind 
edge of eye and hind edge of opercle; mandibular barbel intermediate 

Fig. 2 Lateral view of the head of the allotype of N. tuberigum, MZB 10565. Suborbital flap clearly evident; anterior naris a short tube, valve-like; small 
tubercles are scattered over the head. 
