cribrarius weighing 3,218 g (3,209 from the surf and 9 

 from the tidal pool) were captured. The relationship of 

 mean number per collection and mean station water 

 temperature (in the surf by months) is shown in Figure 1; 

 and carapace width frequency data for tidal pool and surf 

 combined, in Table 18. Ovigerous females were collected 

 as early as mid-June and recently spawned ones as late 

 as mid-September. These data, along with the presence 

 of small individuals (20 mm CW or smaller) from May 

 through October, show that A. cribrarius has a prolonged 

 spawning period. 



Alosa aestivalis (Mitchill) (Clupeidae). In five collec- 

 tions (all from the surf), 22 specimens of 42 to 62 mm SL 

 of Alosa were collected. As Berry (1964) pointed out, it is 

 difficult to identify small specimens of Alosa using any 

 description or key. The characters which appear to be 

 most useful in identifying the species of the Atlantic 

 drainage (A. sapidissima, A. mediocris, A. aestivalis, and 

 A. pseudoharengus) are the number of gill rakers on the 

 lower limb of the anterior gill arch and total number of 

 vertebrae (Hildebrand 1963; Berry 1964). Our specimens 

 have lower-limb gill raker counts of 36 to 41 (x = 37.82) 

 and vertebral counts of 49 to 51 (x = 50.00). The high gill 

 raker counts eliminate Alosa mediocris and the low 

 vertebral counts remove A. sapidissima from further con- 

 sideration. According to Hildebrand (1963), Alosa 

 aestivalis has 49 to 53 vertebrae, whereas A. 

 pseudoharengus has 46 to 50. Both A. aestivalis and A. 

 pseudoharengus show an increase in number of gill rakers 

 with growth until adult sizes are reached (Hildebrand 

 1963). Hildebrand (1963) gave lower-limb gill raker 

 counts as follows: A. aestivalis, specimens of 30 to 49 mm 

 SL— 28 to 36, specimens of 50 to 69 mm SL— 30 to 39; A. 

 pseudoharengus, specimens of 30 to 49 mm SL — 25 to 33, 

 specimens of 50 to 69 mm SL — 32 to 36. Our material, 



then, is more similar to Alosa aestivalis (Mitchill) than 

 to Alosa pseudoharengus (Wilson). We have, therefore, 

 considered our specimens of Alosa as A. aestivalis. 



Dorosoma petenense (Giinther) (Clupeidae). One 

 specimen of 73 mm SL of Dorosoma petenense was 

 collected in the surf on 24 August 1970. Although D. 

 petenense has been introduced into freshwater reservoirs 

 in the southeast, we have found no record published prior 

 to 1974 of its occurring in the estuarine or marine waters 

 of South Carolina. According to Miller (1963), it has 

 been collected from the Gulf Coast of the United States 

 and southward to northern Guatemala and British Hon- 

 duras. Donald C. Scott (University of Georgia) stated 

 (pers. commun.) that in addition to its having been in- 

 troduced into reservoirs in Georgia, there was a native 

 stock of D. petenense present at least as far back as 1955 

 in the Satilla River which drains into the Atlantic, and, 

 at present, it is widespread along the coast having been 

 taken in beach collections at Sapelo and near Brunswick 

 and Savannah. Miller and Jorgenson (1969) reported this 

 species from collections made in the surf at St. Simons 

 Island, Ga. In addition to the specimen collected at Folly 

 Beach, one was collected on 9 February 1970 near Dewees 

 Inlet at the northeast end of the Isle of Palms, in 

 Charleston County (lat. 32°48.8'N, long. 79°43.0'W). 

 Shealy et al. (1974) collected numerous specimens in 

 1973-74 during a survey of South Carolina's estuaries. 



Anchoa mitchilli (Valenciennes) (Engraulidae). This 

 species was collected in all seasons of both years, least 

 commonly in winter. In 30 collections (29 from the surf 

 and 1 from the tidal pool), 372 specimens of 20 to 60 mm 

 SL (362, 20-60 mm SL, from the surf and 10, 47-58 mm 

 SL, from the tidal pool) of A. mitchilli weighing 365 g 

 (350 from the surf and 15 from the tidal pool) were 

 collected. 



Table 18.— Carapace width frequencies (%) by month for Arenaeus cribrarius collected in tidal pool and surf combined during the Folly Beach 

 survey. One specimen of 140.8 mm CW, collected in August 1971, is denoted by an asterisk. 



Month 



Carapace width (mm) 



1-10 



11-20 21-30 31-40 41-50 51-60 61-70 71-80 81-90 91-100 101-110 111-120 121-130 131-140 



N 



1969 

 Oct. 



Nov. 



1970 

 Apr. 

 May 

 June 

 July 

 Aug. 

 Sept. 

 Oct. 

 Nov. 



1971 

 Mar. 

 Apr. 

 May 

 June 

 July 

 Aug. 

 Sept. 

 Oct. 



4.3 



13.0 





34.8 

 100.0 



4.3 

 50.0 



8.7 



17.4 



4.3 

 50.0 



13.0 





12.5 



25.0 



12.5 



12.5 



25.0 



12.5 









1.7 



15.3 



42.4 



23.7 



3.4 



11.9 



1.7 







25.0 



12.5 



25.0 



12.5 



12.5 





12.5 







4.4 



86.8 



8.8 















11.1 



33.3 



44.4 







11.1 









27.3 



27.3 

 100.0 



100.0 



18.2 



9.1 



9.1 





9.1 









41.7 



33.3 



8.3 





8.3 







8.3 





20.0 



6.7 



13.3 





60.0 











46.7 



20.0 







6.7 







20.0 



20.4 



51.0 



10.2 



10.2 



2.0 







4.1 





60.0 



20.0 



5.7 



2.9 



2.9 



2.9 







2.9 



6.5 



50.0 



23.9 



2.2 





2.2 



2.2 



2.2 





3.2 





12.9 



12.9 



35.5 



12.9 



12.9 



6.5 



3.2 



4.3 



6.7 



2.2 



2.2 



2.0 



2.9* 



2.2 



23 



2 



1 



8 

 59 



8 

 91 



9 

 11 



1 



1 



12 

 15 

 15 

 49 

 35 

 46 

 31 

 417 



15 



