92 UPPER CRETACEOUS-LOWER TERTIARY FORAMINIFERA 



Glaessner (1937a) raised Cushman's variety to specific rank, but again gave no 

 figures. He was followed by Cita (1948), Bolli (1951), Noth (1951), Subbotina 

 (1953), Ayala (1954), Nakkady & Osman (1954) and Troelsen (1955), but the figures 

 given by both the first and last authors are different from Cushman's holotype. 



Gandolfi (1955) described two new subspecies of G. contusa (Cushman) and thus 

 changed its name to G. contusa contusa, to distinguish it from G. contusa patelliformis 

 Gandolfi and G. contusa scutilla Gandolfi. Hofker (1956a, 1960a, 1962a) assigned 

 the present from to his genus Marginotruncana which is a junior synonym of Globo- 

 truncana as stated above. Globotruncana conica var. plicata White 1928 is probably 

 a junior synonym of G. contusa contusa (Cushman), but White's brief description does 

 not allow a definite decision without examination of his holotype. Similarly, 

 comparison of oriented thin sections of G. contusa (Cushman) with the holotype 

 of Rosalina linnet mut. caliciforme de Lapparent 1918, and with Globotruncana linnei 

 caliciformis (de Lapparent) of Vogler (1941), showed the possibility that G. contusa 

 (Cushman) 1926 may be a junior synonym of G. caliciformis (de Lapparent) 1918. 

 However, examination of several samples from the type locality of de Lapparent 

 (The Hendaye region of southwestern France) is needed before any decision can be 

 taken, as his original description is very brief, and his figure is only of a thin section. 

 On the other hand, forms described as G. caliciformis by authors are different from 

 the holotype of de Lapparent (1918) and the hypotype of Vogler (1941), and should 

 be renamed and redescribed in more detail. Similarly, G. contusa (Cushman) of 

 Troelsen (1955) is different from the holotype of Cushman (1926), and should also be 

 renamed and redescribed ; the form figured by Berggren (1062) as G. contusa is 

 doubtfully related to Troelsen's form, and is completely different from the holotype 

 of Cushman. 



Globotruncana contusa contusa (Cushman) is believed to have evolved from G. 

 fornicata fornicata Plummer through G. contusa witwickae subsp. nov. as suggested 

 by the morphological characters and stratigraphical ranges of these forms. However 

 Gandolfi (1955) suggested the evolution of G. contusa contusa (Cushman) from 

 G. contusa patelliformis Gandolfi although he admitted its relationship to G. fornicata 

 fornicata Plummer. 



Hypotypes. P.45521. 



Horizon and locality. Figured specimens, from sample No. 18, W. El- 

 Sharawna section. 



Stratigraphical range. Globotruncana contusa contusa (Cushman) was describ- 

 ed from the Maestrichtian Mendez shale of Mexico, and was later recorded from the 

 same formation by Cushman (1927, 1946), White (1928), and Hay (i960). It was 

 also recorded from the Maestrichtian of the U.S.S.R. (Morozova 1939, Keller 1946, 

 Subbotina 1953) ; the Campanian-Maestrichtian of Austria and Switzerland 

 (Noth 1951) ; the Maestrichtian of Trinidad (Bolli 1951, I957«) ; the Maestrichtian 

 of Qabeliat and the Campanian-Maestrichtian of the Sudr sections, Sinai, Egypt 

 (Nakkady & Osman 1954) ; the Maestrichtian Colon shale of northeastern Colombia 



