210 UPPER CRETACEOUS-LOWER TERTIARY FORAMINIFERA 



directly or indirectly evolved, and with which it has been often confused. Bolli 

 (1957&) raised this variety to specific rank, changing its name to G. elongata Glaessner. 

 However, the forms figured by him are different from the holotype of Glaessner, but 

 most probably belong to the present species. He was followed by Loeblich & Tappan 

 (1957a), Olsson (i960) and Hillebrandt (1962). Loeblich & Tappan included in 

 G. elongata forms which are identical with the present species, and others which lack 

 the radially elongated last chamber, characteristic of the species (see synonymy). 

 These forms, as can be seen from their figures, should be considered separately. 

 The form figured by Olsson (i960) as G. elongata Glaessner conforms well with the 

 present species, while that described by Hillebrandt is different from both the holo- 

 type of Glaessner and the present form. 



Globorotalia emilei sp. nov. was confused with G. membranacea (Ehrenberg) 

 and with G. compressa (Plummer) as seen in synonymy. Globorotalia membranacea, 

 although much confused, is a Pliocene form, while G. compressa is only known from 

 the Danian. 



Globorotalia emilei sp. nov. is believed to have evolved from G. compressa (Plum- 

 mer) by the increase in the size of test and the development of the strongly elongated 

 chambers in the last whorl, and the partially developed keel. On the other hand, it 

 is believed to have evolved in two directions : one leading to G. pseudomenardii 

 Bolli, by the development of the marginal keel and the sharply acute axial periphery, 

 and the other to G. troelseni Loeblich & Tappan, by the development of the tendency 

 towards a somewhat evolute coiling in the last whorl. The present species is named 

 after Professor Emile A. Pessagno, Jr., of the University of California, Davis, 

 California. 



Holotype. P.45606. 



Paratypes. P.45668. 



Horizon and locality. Holo- and paratypes, from sample No. 33, Gebel 

 Owaina section. 



Stratigraphical range. Globorotalia emilei sp. nov. ranges throughout the Middle 

 and most of the Upper Paleocene of the Esna-Idfu region. It appears at the base 

 of the G. angulata Zone, and continues as a common to abundant form to the basal 

 part of the G. aequaJG. esnaensis Subzone. Bolli (1957&) recorded his G. elongata 

 Glaessner, which most probably belongs to G. emilei, from the Paleocene, lower 

 Lizard Springs formation of Trinidad, where it was shown to range through the 

 G. pseudomenardii-G. velascoensis Zones. It was also recorded as G. elongata 

 Glaessner, from the Paleocene of the Gulf and Atlantic Coastal plains and of Mexico 

 (Loeblich & Tappan 1957a ; Olsson i960), as G. membranacea (Ehrenberg) from the 

 Paleocene of the Maqfi section, Farafra Oasis, Egypt (Le Roy 1953), and as G. 

 compressa (Plummer) from the Paleocene of various parts of the world (see synonymy). 



Globorotalia esnaensis (Le Roy) 

 (PI. 21, figs. 6a-c) 

 figs. 8-10. 



