36 CALCAREOUS ALGAE OF THE MIDDLE EAST 



Clypeina sp. (Palaeocene) 

 (PL 5. %. 2) 



i960 Clypeina. spp. Elliott: 225. 



Apart from C. merienda, there occur rarely in the Palaeocene-Lower Eocene of the 

 Middle East small Clypeina spp. inviting comparison with those from the French 

 Eocene described by L. and J. Morellet (1913 ; 1922 ; 1939 ; see also Rezak 1957). 

 Random thin sections of these have been noted in material from the Palaeocene 

 Sinjar Limestone of Banik, Mosul Liwa, northern Iraq ; from the Palaeocene 

 Ghurna Beds (Umm er Radhama Formation) of Al Ghurra, Divvaniya Liwa, southern 

 Iraq ; from the Palaeocene-Lower Eocene of Sahil Maleh, Batinah Coast, Oman, 

 Arabia ; and from the Palaeocene of Aqabar Khemer, Hajar, Hadhramaut. The 

 example figured is typical and shows a verticil of 1 mm. diameter, central aperture of 

 0-36 mm. diameter, with about 22 adjacent sporangial tubules. None of these 

 localities have yielded enough material for a precise determination by comparison 

 with the similarly-sized, well-known and beautifully-preserved European material. 



? Clypeina sp. (Permian) 



(PI. 5, figs, i, 3) 



1958a Clypeina Mich, (ou genre nouveau tres voisin) Emberger : 51. 



i960 ^Clypeina sp. Elliott : 219. 



1965 Eoclypeina Emberger MS ; Glinzboeckel and Rabate : pi. 74. 



In a preliminary note on the Upper Permian of Djebel Tebaga, southern Tunisia, 

 Emberger (1958a) listed a clypeiniform alga of which he proposes to describe three 

 new species ; this has been illustrated but not described in Glinzboeckel and Rabate 

 (1965). Debris of the same or a similar form is now figured from the Permian of 

 Iraqi Kurdistan, where it occurs rarely at Harur (Mosul Liwa), both from the base 

 of the Satina Evaporite formation and from the top of the Zinnar Formation 

 immediately below. Whether this is a true Clypeina, ancestral form or homoeo- 

 morph, it seems to represent an early attainment of the umbrella-like sporangial disc 

 familiar in certain Mesozoic and Tertiary genera : it is hoped that M. Emberger's 

 descriptions will throw light on this. The Jugoslav Permian dasyclads Salopekiella 

 and Likanella (Milanovic 1965 ; 1966) bear no close resemblance, and nothing else 

 associated is at all comparable. 



The recent description of Clypeina besici Pantic (1965) from the Upper Triassic of 

 Jugoslavia is a valuable confirmatory link between the Permian Eoclypeina and the 

 familiar Upper Jurassic C. jurassica. 



Genus CYLINDROPORELLA Johnson 



Diagnosis. Cylindrical calcareous bodies terminally tapered or rounded, 

 interpreted as serial dasyclad units arranged in life somewhat similarly to those of the 

 Recent Cymopolia. Internally the longitudinal central canal (stem-cell cavity) is 

 surrounded by rings of proportionally large spherical sporangial cavities alternating 



