78 JURASSIC RHYNCHONELLIDS 
RemARKs. The form was first mentioned by Buckman and Walker in 1889 in 
their discussion of the spinose Rhynchonellidae, when they described it under the 
name of Acanthothyris senticosa var. fileyensis and suggested that it resembled A. 
senticosa of Orbigny. However, as A. (Echinirhynchia) fileyensis shows marked 
differences from both the specimens in the Orbigny Collection and A. (Echinir- 
hynchia) senticosa (Schlotheim), it has been decided to describe it as separate species 
despite the paucity of material. 
Subfamily TETRARHYNCHIINAE Ager, 1965) 
Genus SOMALIRHYNCHIA Weir 
1925 Somalivhynchia Weir: 79. 
1929 Somalirhynchia Weir; Weir: 38-39. 
1935 Somalivhynchia Weir; Muir-Wood: 93. 
1964 Praecyclothyris Makridin, 150-51. 
1965b Somalivhynchia Weir; Ager: H614. 
TyPE sPEcIES. S. africana Weir, by original designation. 
EMENDED DIAGNOsIs. Large, subpentagonal, trilobate rhynchonellids ; 20-30 
coarse, simple, subangular ribs ; large, sub-erect beak ; possesses septalial plates 
and median septum, which together form a septalium ; crura radulifer. 
STRATIGRAPHICAL RANGE. Upper Oxfordian—Lower Kimmeridgian. 
DISTRIBUTION. Somaliland, Syria, N.W. Europe, Russia. 
REMARKS. Weir (1925, 1929) gave the name S. africana to the species figured by 
Noetling (1886) as Rhynchonella moravica Uhlig ; he suggested that the differences in 
outline and particularly in muscle scar patterns necessitated regarding R. moravica 
Noetling as differing from R. moravica Uhlig not only specifically but generically. 
The author considers that Weir was probably correct in establishing a new species 
for Noetling’s R. moravica. However, in internal structure, as well as general 
external appearance, S. africana, in the author’s opinion, closely resembles R. 
moravica Uhlig and should be regarded as congeneric with it. 
In basing his taxonomy almost entirely on the muscle scar patterns, Weir was 
closely following the ideas of Buckman. However, it is now considered that the 
similarity exhibited by all the other features outweigh in taxonomic significance the 
dissimilarity of the muscle patterns. The internal details of R. moravica Uhlig were 
figured by Wisniewska (1932) and, allowing for the differences in appearance caused 
by her angle of sectioning, are closely comparable to those given by Muir-Wood 
(1935) for S. africana. They appear to have little in common with Septaliphoria 
arduennensis (Oppel), the type species of Septaliphoria Leidhold, to which genus R. 
moravica Uhlig was assigned by Wisniewska. Consequently, the author does not 
understand on what grounds Muir-Wood (1935) remarked, ‘‘ Further study of the 
internal characters of R. moravica by Wisniewska (1932, p. 22) has emphasized the 
differences between the two species (i.e. R. moravica Uhlig and S. africana Weir). 
R. moravica is now assigned to the genus Septaliphoria (Leidhold 1920) ”’. 
The wide geographical distribution of the genus is coupled with a very limited 
stratigraphical range. It is considered that the distribution of the genus can be 
