404 DR MORRISON WATSON AND MR ALFRED H. YOUNG 
respect, therefore, Beluga agrees with these. The description of the form of 
the stomach in different species is not sufficiently exact to enable us to come 
to any conclusion with regard to the relation in which the latter stand to one 
another; but founding upon Dr Murte’s* observation that the second and 
fourth compartments of the stomach of Risso’s Grampus differ from those of 
Globiocephalus in respect of their more elongated and less globular form in the 
- former, we may state that so far as this organ is concerned, Beluga appears to 
be more closely related to Grampus than to Globiocephalus. | 
In Grampus Rissoanus, however, Dr Murte observes that the opening, by 
means of which the second communicates with the third stomach, is situated 
close to that between the first and second compartments; whereas in Beluga, as 
we have seen, they are separated by an interval equal to two-thirds of the length 
of the second stomach. In this respect Beluga agrees with G'lobiocephalus 
rather than Grampus. 
We have previously directed attention to the presence of a fold of mucous 
membrane, extending between the cesophageal aperture of the first and the 
superior wall of the second compartment of the stomach in Beluga, and 
remarked upon the resemblance which it bears to the superior lip of the 
cesophageal groove of the ruminant stomach. A similar observation has been 
made by Dr Murte in his description of Globiocephalus. Whilst directing 
attention to the presence of this fold, however, we would wish to avoid attri- 
buting to it any physiological significance in tracing an analogy between the 
stomach of the ruminants and that of the cetaceans. Professor TuRNER,t 
founding on his observation that in Globiocephalus the cesophagus communicates 
freely with both the first and second stomachs, is of opinion “that a provision 
would seem to exist in this animal for permitting a process of rumination as far 
as regards the contents of these two compartments, and an additional link is 
established between the ruminant and cetacean stomach.” 
With this opinion we are unable to agree—-Firstly, Because im the rumi- 
nants, in which alone do we know anything positive regarding the process of 
rumination, the food, after passing through the first, is regurgitated from the 
second stomach, and subsequent to undergoing a second process of mastication 
is passed into the zhird stomach, this transference of the bolus being effected by 
means of the cesophageal groove. In the cetacea, on the other hand, at least in 
Beluga and Globiocephalus, the cesophageal groove is incomplete, and is there- 
fore wholly inadequate to perform the function of a canal, by means of which 
the food could be transferred from one compartment of the stomach to another. _ 
It, moreover, differs from the corresponding structure in the ruminant stomach, = 
inasmuch as it terminates in the second compartment of the viscus instead of 
* X. p. 132. + IX. p. 72, 
