408 DR MORRISON WATSON AND MR ALFRED H. YOUNG 
septal processes pass inward to the hepatic substance. There is no trace of a 
gall bladder. Neither did we observe any dilatation of the hepatic duct in the 
wall of the intestine similar to that described by Dr Murte* in Globiocephalus. 
The hepatic duct measures 5 inches in length, and is united with the pancreatic 
duct an inch and a half before it reaches the wall of the intestine. The com- 
bined ducts enter the duodenum 4 inches beyond the pylorus. 
In respect of the absence of any trace of subdivision of the liver into a right 
and left lobe, Beluga appears to differ from almost every cetacean in which that 
organ has hitherto been described. That this isnot an exceptional arrangement, 
occurring in only a single specimen of the species, appears to be proved by the 
fact that neither Barctay and NEILL nor WYMAN make any mention of distinct 
hepatic lobes in the specimens which they examined; at the same time, it 
should be observed that variation in the relative sizes of the hepatic lobes in 
different individuals of the same species of cetacea does sometimes occur, as is 
evident from the observations of Jackson and Murie upon Globiocephalus, in 
which, according to the former, the right is two or three times larger than the left 
hepatic lobe, whilst the latter investigator found them to be of the same size. — 
On the other hand, Beluga agrees with all the other cetacea (with the exception 
of Globiocephalus chinensis)t in the absence of a gall bladder, and with the 
majority of the toothed whales,{ in respect of the junction of the hepatic and 
pancreatic ducts previous to their passage through the intestinal wall. In 
Globiocephalus§ and G'rampus || Dr Murte describes the hepatic duct as being 
dilated into a kind of reservoir, where it lies in the wall of the duodenum. No 
such arrangement was distinguishable in Beluga. 
Pancreas.—This organ measures 10 inches in length and 23 in greatest 
breadth. It is somewhat flattened from above downward, and its extremities 
are evenly rounded. It extends from the right side of the apex of the first 
gastric compartment to the pylorus, and is overlapped on its lower surface by 
the second cavity of the stomach. Its surface is distinctly lobulated. The 
duct, as already stated, unites with that of the liver an inch and a half before 
the latter passes through the intestinal wall. 
The pancreas does not present much variety either in respect of form or 
relations in the cetacea. In Beluga it does not appear to differ in either of 
these particulars from what has been noticed in the other toothed whales. 
Spleen.—This organ is small and cake-like. It is situated directly behind 
and in contact with the superior surface of the first gastric compartment, and 
touches the left extremity of the pancreas. It is irregularly rounded, and 
measures about 3 inches in diameter, with an average thickness of 1 inch. 
T. p. 261. { IL p. 261. 
az ll 
¢ (In Zyphiorrhynchus these ducts open separately into the duodenum.) 
§ IL p. 261. |X. p. 133. 
* 
