12 TRANSACTIONS LIVERPOOL BIOLOGICAL SOCIETY. 
placed in a cage outside for the night, but in the morning 
it was found that all the females had disappeared, they 
having made a tunnel through the floor of their prison, 
and having thus got away, whilst the three males remained 
in the cage. In the breeding season, also, the adult 
females were all found to have their claws blunted as the 
result of their digging operations, whilst the claws of the 
males were perfectly sharp. On the other hand, the 
males, as in other Struthious birds, appear to do all the 
duty of incubation, and also take entire charge of the young 
till they are able to shift for themselves. So that the two 
sexes of this bird appear to have established a well-defined 
division of labour with regard to their nesting operations. 
It is said, indeed, by the natives, that after laying her two 
eggs in the nesting cavity prepared by herself, the female 
goes off and pairs with another male, forming a new nest 
elsewhere. 
The wings are extremely aborted, and the condition of 
these and the parts of the skeleton connected with them, 
is of great interest. 
To begin with, a distinct alar membrane is present. In 
the wing of A. bullerz there is clearly manifest a definite 
fold of skin or pre-patagium passing between the upper- 
arm and the fore-arm, and an equally well-marked fold or 
post-patagium between the upper-arm and the trunk; 
this is precisely what we meet with in typical flying birds, 
and it is very suggestive. Furthermore, there are clear 
remains of remiges or flight feathers in the wings. Thus, 
T. J. Parker found 8 to 4 metacarpo-digitals and 9 to 10 
cubitals in the fresh wing of A. bullevz, and on the outer 
side of the wing there was a distinct row of tectrices. 
majores, or greater wing coverts. Though so small and 
reduced, these remains of flight feathers point definitely 
to a time when they served the purpose for which flight 
