212 DR J. 3J. MACFARLANE ON THE 



1. Lapageria proportionately in depth as 4 to 3 in Lapageria. Each cell is tangentially flattened, 

 -• Phiiageria an d the cuticular layer is continued inwards in wedge fashion along the lateral walls. 



Veitchii. J . 



s. Phiiesia buxi- Its average size is 100 ft long by 30 ft wide and 25 ft deep. In 2 the free epidermal cell 

 faces have ridges 2^ to 3 ft deep (figs. 2, 5). Each cell is intermediate in shape, and 

 measures 80 ft long by 30 ft wide and 35 ft deep. It should be said, however, that while 

 this is the average of many measurements, the cells are variable in size. The cuticle 

 dips in along the lateral walls in wedge-like fashion as in 3. 



The outer cortex in 1 is made up (fig. 3) of twenty to twenty-five layers of large, 

 moderately thickened cells, which pass abruptly into an inner cortex of fifteen to twenty 

 layers of dense sclerenchymatous cells nearly or quite uniform in thickening and trans- 

 lucency. Each cell wall of the latter elements shows four to five thickening lamellae. 

 In 3 (fig. 1 ) the outer cortex consists of nine to ten layers, the cells of which are larger and less 

 thickened than in the last. The inner cortex shows three to four indurated layers, of which 

 one or two external are greatly thickened and brown pigmented, the internal ones having 

 clear and less thickened walls. The former exhibit eleven or twelve thickening lamellae. In 

 2 (fig. 2) there are fifteen to seventeen layers in the outer cortex, and in the inner nine to 

 eleven. The outermost cells of the latter are slightly pigmented brown if mature stems 

 are chosen near the level of the ground. Each wall has seven to nine thickening lamellae. 



Longitudinal views of the three demonstrate that there is a very pretty intermediate 

 condition in the hybrid between the numerous and distinct wall pits of the outer cortex 

 in 1, and the few and faintly-marked pits in 3. 



The central parenchyma of the stem is small-celled, thick-walled, and pretty uniform 

 in 1 ; that of 3 is large-celled, thin-walled, except for a few isolated and more strongly 

 indurated elements scattered irregularly. The hybrid is very closely between these 

 conditions. In longitudinal view the relative amount of thickening and distribution of 

 the wall pits is equally noticeable here as in the outer cortex. 



Vascular Bundle System. — The stem bundles appear in all cases to be greatly larger 

 in 1 than in the other parent or hybrid, the average diameters being as 400 ft in 1 to 

 250 ft in 2, and 180 to 200 ft in 3. If, therefore, the hybrid material was sufficiently 

 matured, it approaches here to the pollen parent. 



In 1 the phloem patches (fig. 9) are 85 to 90 ft deep, and are made up of large sieve- 

 tubes, each 40 to 45 ft across, along with others that are smaller but of varying size ; also 

 of companion cells 8 to 10 ft across. The xylem has a flat or slightly convex face next the 

 stem centre, and its main mass is made up of two large scalariform vasa. # Each vas is 100 

 to 120 ft across, and between each pair are smaller radially- elongated scalariform or pitted 

 vasa (or tracheids), and in line with the front of the vasa is a small protoxylem patch. 



In 3 the phloem patches (fig. 7) are 45 to 50 ft deep, the sieve-tubes are nearly uniform 

 in size, unlike those of 1, and measure 20 to 25 ft across, while the companion cells are 5 to 

 8 ft. The xylem is typically wedge-shaped, the back part next the phloem being occupied 



* I ii compound or branching bundles several of these, usually of smaller size, may represent the above, and the same 

 is true regarding the others treated of. 



