248 DR J. M. MACFARLANE ON THE 



' C Sprcerian™ m abidance of gland-tipped ones derived from the first parent. They are, however, less 

 -'• Cypnpedium abundant, for over equal areas under Zeiss' A objective, there were sixteen to eighteen 



Leeanum. ~ J ' & 



5. cypripedium m Q Spicevianum as compared with nine to eleven in the hybrid. The spotted areas 



insigne. ± L •> r 



in the hybrid are quite glabrous, as in the other parent. This mode of hair distribution, 

 alike as to position and relation to epidermal colour distribution, is all the more remark- 

 able when we remember that the gland hairs of C. Spicevianum and the hybrid spring 

 from colourless cells, but are nevertheless filled, in at least the three to four lowest, 

 with a pigment like that filling the epidermal cells, which form the areas that are invari- 

 ably glabrous in C. insigne and in the hybrid. Further, while along the dark purple 

 line that traverses the middle of the sepal in C. Spicevianum, both types of hair of a 

 ruby colour spring from similarly coloured cells, and while the middle of the sepal in C. 

 insigne is glabrous and colourless, the purple line is reproduced in the hybrid though in 

 a rather diffused state, and both kinds of hair spring from ruby cells of it. 



The median inferior (in position) sepal shows in C. Spicevianum a moderate number 

 of simple multicellular hairs externally, and great wealth of glandular hairs internally. 

 C. insigne shows the converse condition, viz., abundant glandular hairs externally, and 

 fewer simple hairs internally, though the relative numbers are more nearly equal than in 

 the former. In the hybrid there is a pretty uniform distribution of both types alike 

 on the outer and inner sides. 



Petals. — The lateral petals are mainly of interest from their hair distribution, but 

 they may be passed over as they do not share the striking peculiarities of the sepals. 

 The outer surface of the labellum has wavy-walled cells, which, for shape, size, &c, are 

 quite alike, though the colour contents vary. Internally they agree, except in colour and 

 hair distribution. C. Spicevianum has hairs spread uniformly along the bottom of the 

 slipper ; C. insigne shows two types, one simple and multicellular, the other short and 

 club-shaped, being composed of three small stalk cells, and a terminal knob cell. In the 

 hybrid both kinds of hair are encountered though reduced in size and number. 



Stamens. — We may neglect the abortive stamens except the crescentic one which 

 represents the third of the outer whorl, and which merits detailed study. In C. 

 Spicevianum, both outer (anterior) and inner (posterior) surfaces of it are quite smooth, 

 and only a very few hairs exist towards the top. It measures from 3 m. at its thinnest 

 to 6*5 m. at its thickest part. In C. insigne, the outer surface is covered by yellowish 

 glassy warts, which appear under the low power of the microscope as magnificent greenish 

 yellow papillae, each ending in a hair. The inner surface is also hair-covered over its 

 upper region. The microscopic resemblances of the hybrid seem all to be towards the 

 latter parent till sections are made and examined, when the size of the papillae and of the 

 cells forming them, as also the hair distribution, are found to be a reduction by half of 

 the conditions of the latter parent. 



The pollen of C. Spicevianum is slightly smaller than that of C. insigne, and is pale 

 yellow, while in the latter it is greenish yellow, and in the hybrid dull yellow. The 

 pollen of the hybrid has a round but rather granular and hard look. 



