1893.] The Ancylopoda, Chalicotherinm and Artionyz. 119 
tarsals, the structure of the ankle and wrist joints, and the 
terminal facets of the metapodials all indicate that the feet 
were sub-digitigrade. Displacement is very rare in planti- 
grade types. The proximal phalanges are peculiar in having 
the metapodial facets directed obliquely upward; both these- 
and the median phalanges are short; the distal phalanges are 
` very peculiar, they present a deep terminal cleft and were evi- 
dently drawn up or retracted, as in the cats, instead of being 
drawn down or flexed, as in the Edentates. Thus, it is proba- 
ble that Chalicotherium walked more like a clumsy-footed cat 
than like the bear. The soles of the feet were slightly turned 
inward and the fore-limb was adapted to digging or partial 
prehension. : 
Restoration. The axial skeleton in the accompanying draw- 
ing is mainly based upon Filhol’s restoration, but the scapula 
and pelvis are independently restored upon the perissodactyl 
model; the sloping trunk leads us to expect a rather broad 
` pelvic basin. All the above parts are purely conjectural. 
The proportions of the fore and hind limbs with each other 
and with the skull are based upon the figures of Depéret. In 
Filhol’s drawing the skull is proportionately too small. The 
side views of the feet are after Filhol’s excellent drawings, but 
the articular positions are drawn differently, for M. Filhol 
makes the animal fully plantigrade, while the metapodial and 
phalangeal and higher articulations prove that it was digiti- 
grade. s 
What is the explanation of this remarkable assemblage of 
characters? Where shall we place Chalicotherium, with ref- 
erence to the two great orders between which it seems to stand 
midway. If, observing the teeth and the carpus and tarsus, 
we relate it to Perissodactyla, we find ourselves placing a 
clawed type among hoofed types. Or is it to be considered an 
Edentate because of its claws and edentulous premaxillaries 
as Filhol has suggested? Or is it representative of a distinct 
order midway between the Ungulata and Unguiculata? It is 
too early to attempt a final solution of these questions. We 
must wait for the discovery of the middle and upper Eocene 
Chalicotheriide. It is evident from Gaudry’s discovery of the 
