ELEDONE. 495 
From the above description it can be seen that, 
although water may enter the mantle cavity all along the 
external opening of the mantle, yet when the funnel is 
locked the only way out for the contents of the mantle 
cavity—excretory or genital products, water, &c.—is 
through the central funnel chamber. _ As in other 
Octopods, Eledone has no valve in the funnel. It has, 
however, a large and elaborate mucous gland—Miiller’s 
eland (Pl. II, fig. 10, f.0.). This is four-lobed, and is an 
elaboration of the internal epithelium of the funnel, and 
may best be seen by opening the funnel ventrally, as in 
Pl. II, fig. 15. It serves to lubricate the internal surface 
of the funnel, which consequently is generally coated over 
with opaline viscous mucus, rendering the gland itself 
rather obscure in fresh specimens. It may be seen, how- 
ever, on scraping the mucus away. 
Pallial Complex (figs. 8 and 11).—Under this general 
term may be included those important organs situated in 
the mantle cavity, together with the external apertures 
of certain internal organs. Eledone, like other Cephalo- 
pods, in spite of its high specialisation along certain lines, 
has yet retained its primitive symmetry in certain 
features, including the pallial complex. The organs of 
the pallial complex are :— 
(1) A pair of Gills, one on each side of the visceral 
mass, and attached to it by muscles, vessels, &c. 
(fig. 11, g.); 
(2) The Anus, situated anteriorly, between the left 
and right halves of the vertical septum (fig. 11, an.) ; 
(3) The Urinary papillae—one pair, protruding for 
about 12 mm. in a large specimen into the mantle cavity, 
just in the angle between the base of the gill and the 
visceral mass (fig. 11, Ur. p.). The urinary aperture is a 
smal] hole at the tip of this papilla; 
