304 The American Naturalist. [April, 
color of the leaflets in the native plant being light lively green, 
with little difference between the two surfaces. In these 
points of difference, too, the garden berries are characteristic- 
ally like the Chilian. The truss or inflorescence is different 
in the two. In the garden berries, the truss stands more or 
less oblique or is often prostrate, and it is broken up into two 
or three strong, often unequal spreading arms from which the 
short and stout fruit-stems spring, and this is the distinctive 
habit of the Chilian species; in the Illinoensis, the truss is erect 
and it breaks up more regularly at its top and the inflorescence 
is less strongly spreading in proportion to the number of fruits 
it contains, and the fruit-stems are weak and slender and more 
or less drooping. The calyx is very large in the garden ber- 
ries, a fact which Duchesne recorded in the name’ Fragaria 
calyculata which he applied to the large-hulled forms like the 
old Bath Scarlet, of which many are in cultivation at the pres- 
enttime. The fruit in Illinoensis is small and soft and bright 
scarlet, usually with a distinct neck and deeply embedded 
seeds; that of the garden berries still maintains the features 
of the Chilian berry in its large size, mostly globular-pointed 
form, dark color and seeds borne more nearly upon the 
surface. The garden berries are in every way much farther 
removed from the native berry than they are from the Chilian. 
From the latter they differ most widely, as I have said, in the 
taller growth and less hairiness;" but even in these features 
they do not resemble very closely the Jllinoensis. It may be 
urged that all these differences might have come about under 
the influence of cultivation if Illinoensis itself had been the 
parent of the garden forms, to which I reply that direct 
experiment does not sustain the assum ption, and that the excel- 
lent engravings of the early forms of the Pine strawberry 
show the same differences. It was the study of these pictures 
which first led me seriously to doubt the East-American origin 
"Itis often said that the fruit of the Chilian strawberry is erect and that the garden 
berries differ in a nodding fruit, but this is an error. While the fruit stems of the 
ï have grown, the fruit has the same drooping habit as in the garden berries. 
Chilian species probably varies naturally in its fruiting habit, but I have yet to rep 
instance in which it holds its fruit upright. 
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