sept. i9io.] SUZUKI.— ON THE STRUCTURE AND AFFINITIES 



193 



of the spherical body parts varies from 50-180 p. The wall 

 of the fructification is made up of thin walled plectenchyma 



(Text-fig. 2 and Phot. 6, 

 PI. VII), and is usually 

 5-7 cells thick, although 

 they can be much thicker 

 or thinner. The orifice of 

 the spherical or flask- 

 shaped fructification is 

 not much prolonged or 

 protruded like a beak, 

 thus the neck of the 

 fructification is short, and 

 plectenchymatous tissue of 

 the orifice is often found 

 underlining the guard cells 

 of the stomates. 



No stroma is develop- 

 ed in connection with 

 the fructifications. In one 

 case a few club-shaped transparent bodies are seen standing 

 from the bottom of the hollow short necked flask (Text-fig. 3). 

 They are naturally seen as the remains of some of the asci, 

 and the fructification as the perithecium. Besides asci, there 

 are a number of thinner bodies seen standing with them. 

 They are interpreted for the present as paraphyses. Un- 

 fortunately neither spores nor any further structure of asci 

 can be seen. 



The spores had been discharged before the asci became 

 fossilized or they might have not yet been fully developed, as 

 the asci we:e still young. 



Affinities. The growing mycel in the parenchyma of the 

 host, the well developed septate hyphae, and the formation of 

 the perithecium locate the fungus among Ascomycetes. The peri- 

 thecium imbedded in the mesophyll and opening outside by an 

 orifice, the spherical or short necked flask-like form of the peri- 

 thecium indicate the association of this fungus with Sphasriales 



Text-fig. 3. — Pleosporiles Shirainus gen. et sp. 

 nov. Perithecium cut longitudinally through the 

 neck and orifice, showing asci (as). pr t paraphyses; 

 hy, hyphae; ep f espidermis of the leaf of host. 

 X370. 



