On the Coupling of Two Leaf=Characters 

 in the Japanese Morning Glory. 1 



By 



Tokio Hagiwara. 



Wmle studying hereditary behavior in the Japanese Morning Glory, 

 I have met with a case of coupling which takes place between two 

 leaf liaracters. Although the study is not yet completed 1 shall publish 

 the results so far obtained as a preliminary report. 



In 1917 crosses were made between three homozygous races, the 

 one, designated as (5) in the tables given below, having variegated 

 and rolled leaves, 2 while the leaves of the other two, designated as 

 (9 A) and (AX), being homogeneously colored and smooth and flat. 

 The Fi plants thus raised were found to have all homogeneous and 

 normal leaves, showing that the variegated and rolled conditions are 

 both recessive characters. 



The variegation in the Morning Glory has already been studied by 

 Takezaki. 3 According to him, the variegation behaves as a simple 

 Mendelian recessive to the normal. Similar results were obtained in 

 my experiments ; the segregating numbers observed in the F* genera- 

 tion being : — 



Observed Expected 



Homogeneously- 

 colored 



Variegated 



Total 



Homogeneously- 

 colored 



Variegated 



. D. 



S.E. 



5x9A —a 12 



4 



16 



12.00 



4.00 



±0.00 



±1.73 



„ — b 187 



67 



254 



190.50 



63.50 



±3.50 



±6.91 



„ -c 10 



4 



14 



10.50 



3.50 



±0.50 



±1.61 



„ -d 34 



10 



44 



33.00 



11.00 



±1.00 



±2.87 



5xAX— a 23 



8 



31 



23.25 



7.75 



±0.25 



±2.41 



„ -b 12 



3 



15 



11.25 



3.75 



±0.75 



±1.67 



Totals 278 



96 



374 



280.50 



93.50 



±2.50 



±8\37 



1. The substance of this paper was published in " Journal of the Scientific 

 Agricultural Society" No. 206, Oct. 1916 (in Japanese). 



2. The margin of the leaves more or less roll upwards, and the surface is usually 

 rather uneven especially in the young lower leaves. 



3. Nippon lkusvugakukai Kaiho (Journal of the Japanese Breeders' Association). 

 Vol. 1, No. 1, 1916." 



