V 



.J 1 . 



\\>rary of Co^ 

 MAR8 791.9 



^ 



^n'an De9^ 



On the Xenia in the Barley. 



By 



3Iasao So and Yoshitaka Imai. 



From tlie data on the inheritance of grain color in barley, Miyazawa 1 

 formulated a hypothesis of vegetative segregation of gene which he 

 a:-Mimcd to have occurred just before the formation of the growing- 

 point of the ear. But the writers' studies treating the same subject 

 have led to quite a different conclusion, namely ; the characters in 

 question arc transmitted by xenia, and are not a sort of vegetative 

 segregation occurring on the ear of the hybrid. So it may be of in- 

 terest to report these results, though they are not yet completed. 



Miyazawa 2 ' thought that the grain color in question is due to the 

 pigment which is present in the cells of pericarp and of testa respective- 

 ly, so there would be no relation to the double fertilization. Actually 

 the pigment,"' however, is present in the aleurone cells which form the 

 outer layers of the endosjjcrm, so the expectation of the effect of the 

 secondary nucleus of pollen-tube may be natural. In fact, such in- 

 fluence w-as experimentally recognized as follows: When the blue grained 



1) 0b< r die hi leaikartige Spaltnng eines Qerstenbastards. Botanical Magazine, 

 Vol. XXX, 1910. li Japanese paper, Ibid, Vol. XXXI, L917. 



II writing went as follows: ' Sie riilirt, wie die anatomiBche Untersuchung 

 inir geseigt bat, eon den in der Frucht-resp. Samenschule rorhandenen V-.w 



rn 1 1 • r . da bei Bekilori die letztere schwarz und viel reichlicber rertrelen Bind, 

 Idenmelon, wobei Bi'e olivengr&n Bind. Ea mag noch besonders erwShni 

 irerden, da i entlicbe Endosperm gans falblos i I and mit < I ■ r Cornerfarbe 



nicliti zii tun ! 



Bbowh already recognized the existence of blue pigment in alenrone cells 

 itilized ih'-, condition in I » • — «. works on the i of eed of barley On the 



ice of a » ■ membrane i i of I In- ( trami- 



neae. Ai al of B itany, Vol. .\.\ I. 190] 



