676 PROFESSOR FRANK J. COLE 



between the cornual cartilage and the lateral labial here, but says nothing as to its 

 nature. Neumayer did not find it at all. Parker and Ayers and Jackson figure a 

 slight projection on the cornual cartilage, and this, I believe, represents the place where 

 the projection was torn off in cleaning up the cornual cartilage, as happened in my own 

 case. Pollard (18), in constructing wax models from serial sections, was bound to see 

 it, and we consequently find it in his fig. 1 1, but he makes no reference to this feature 

 in his description. It may possibly represent the remains of a cartilaginous connection 

 between the cornual cartilage and the lateral labial. 



The nasal skeleton is perhaps chiefly peculiar on account of the structure of the 

 posterior transverse bar of the olfactory capsule (p. t. b.). This sends downwards and 

 backwards a cylindrical rod {h. p.'), which I formerly believed (7, p. 768) was a 

 connection between the nasal capsule and the hypophysial plate. And so it is ; but it 

 is something more. When studying the velo-quadratus muscle my attention was 

 drawn to this rod again (8, p. 721), and the origin of the muscle at once suggested that 

 the rod is really a connection between the nasal capsule and the trabecula, and that 

 it merely fuses with the hypophysial plate on its way to the trabecula. This is a 

 typical illustration of how a dissection, even if carefully carried out, just fails to result 

 in that refined degree of accuracy necessary for the right conclusion. A comparison 

 of fig. 2 of my first part and fig. 2 of the present one will illustrate this point. 

 Neumayer's fig. 4 should also be consulted. 



The nasal capsule varies in histological structure. It is really a mixture of soft and 

 hard cartilage, and either may predominate perhaps according to the age of the animal. 

 The large lateral plate, however (I. p.), always contains more hard cartilage than any 

 other part of the capsule. 



As previously described, the lateral labial cartilage (/. /. c.) exhibits a very curious 

 sigmoid twist before fusing with the basal plate. I never realised, however, how 

 considerable this twist was until I plotted fig. 2. It cannot be shown in a lateral view. 

 Pollard does not appear to have noticed it, and Parker missed it in his dissections, 

 but it appears in his sections as the "oral barbel" — which is further forward than 

 where he figures it, nor could it have been in that position. Similarly, it is not 

 described by J. Muller and Ayers and Jackson. The only writer who has noticed it 

 is Neumayer, in whose fig. 4 this mysterious twist is even more marked than in my 

 fig. 2. Its meaning I am unable to explain. 



In the neighbourhood of the sigmoid twist on each side and embedded in the soft 

 pseudo-cartilage at the anterior end of the basal plate is a discrete piece of soft 

 cartilage (figs. 1 and 2). The two pieces form a distinct pair (cp. fig. 2), and appear 

 to have some significance. 



The cartilage of the fourth tentacle (4) consists of both the hard and soft varieties. 

 The former is confined to that part of the cartilage situated within the contour of the 

 body (the internal rod), and in the old specimen on which my former figures were 

 based this rod was formed largely of hard cartilage. In the present specimen, however, 



