356 MR R. C. DAVIE ON 



Hymenophyllacese it occurs in Hymenophyllum demissum, H. dilatatum (Forst.) 

 Sw., Trichomanes elegans, Rich., and T. radicans, Sw. 



A divergence into the marginal type occurs in Loxsoma Cunning hami, described 

 above, while the type of Balantium culcita is repeated among the Cyatheaceae in 

 Dicksonia fibrosa, Col. In the other members of the Cyatheaceae examined the 

 extramarginal type prevails. These include Cibotium Schiedei, Schlecht. et Cham., 

 Thyrsopteris elegans, Kze., Cyathea mexicana, Schlecht. et Cham., C pubescens, 

 Mett., Hemitelia grandifolia (Willd.) Spr., and Alsophila glauca (Bl.) J. Sm. 



Passing into the Ferns on the border-line between Gradatse and Mixtae, we have 

 the broken leaf-trace in Peranema cyatheoides, Don, and Diacalpe aspidioides, BL, 

 the unbroken leaf-trace in Woodsia ilvensis (L.) P. Br., W. polystichoides, Eat., 

 Matteucia orientalis (Hk.) Trev., M. strutliiopteris (L.) Todaro, and Onoclea sensi- 

 bilis, L. (in the last three the leaf-trace is binary at its departure from the stalk, but 

 unbroken below the pinnae). In all of these the pinna-trace goes off extramarginally 

 from the back of the hook, whether in the unbroken trace or in the adaxial strand of 

 the broken type. 



- In the genus Cystopteris there is a variation ; in Cystopteris fragilis (L.) Bernh. 

 and C. montana (Lam.) Bernh., the supply to the pinnae is marginal. The leaf-traces 

 have slightly hooked ends to each portion of the binary strands. The whole set of 

 tracheides lengthens, a "thickening" of the number occurs about the centre of the 

 leaf-trace on the adaxial face, a break takes place there in the series, and the 

 " daughter " trace separates off. Below the second pair of pinnae the binary trace 

 becomes an unbroken strand, and the supply is there distinctly marginal. The supply 

 to the basal pinnae resembles the type already described for some species of Gymno- 

 gramma and Ceropteris (p. 352), though the thickening is less marked in Cystopteris. 



In the Aspidieae, the leaf-traces are of the broken type and the supply to the 

 pinnae is extramarginal. In Dryopteris phegopteris (L.) C. Chr. there is a gap 

 opposite to the pinna. In Dryopteris pulninulifera (Bedd.) 0. Ktze., D. filix mas 

 (L.) Schott, D. setigera (Bl.) 0. Ktze., D. serrata (Cav.) C. Chr., Didymochlsena 

 truncatula, Polystichum aculeatum (L.) Schott, var. angulare, Pr., P. Standishii 

 (Moore) C. Chr., the supply is extramarginal. In Aspidium Moorei (see above, p. 352), 

 the pinna-trace comes partly, marginally, from the tip of the adaxial strand, and 

 partly from the median strand of the leaf-trace. 



From the Aspidieae forwards into the main mixed Ferns both extramarginal and 

 marginal types of pinna-supply occur. The Davallieae carry forward the extra- 

 marginal type through Leptolepia novsB-zelandise (Col.) Kuhn, Microlepia hirta (Klf.) 

 Pr., M. hirta, var. cristata, M. speluncse (L.) Moore, M. platyphylla (Don) J. Sm., 

 M. strigosa (Thbg.) Pr., and Dennstaedtia adiantoides (H. B. Willd.) Moore. Micro- 

 lepia hirsnta (.J. Sm.) Pr., Nephrolepis Amerpoldii, hort., N. Fosteri, hort., Hill, 

 N. Piersoni, hort., and N. Scottii, hort., all distinctly show the marginal type. 

 Humata repens (L. fil.) Dicls, var. alpina, and Lindsaya repens (Bory) Bedd., are 



