UNITED STATES DEPARTMENT OF AGRICULTURE 
mrs BULLETIN No. 737 | : 
Contribution from the Bureau of Entomology 
L. O. HOWARD, Chief 
4 | Washington, D. C. PROFESSIONAL PAPER March 17, 1919 
# 
‘THE TOBACCO BEETLE: AN IMPORTANT 
PEST IN TOBACCO PRODUCTS 
By 
G. A. RUNNER 
Entomological Assistant 
[With Technical Descriptions of Coleopterous larve 
By Adam G, Boving ] 
CONTENTS 
; Pago 
A Pest in Cured and Manufactured To- Description of Stages) . . 2 2» « « «% 412 
Economic History . . . 1 2 6 e 
bacco; ItsCommon Names ... . 1 | Life Historyand Habits ..... - 14 
The Character of lisInjury ..... 2 | Seasonal History . . « « « « «© «© « 20 
Classification and Synonymy. . .. - 4 | Insects Likely to be Mistaken for the x 
_Food Substances of the Tobacco Beetle. § 5 Tobacco Beetle . . . 6 » « « » - 27 Hl 
Food Habits of Beetles Related to the Natural Control. . 2. 2 « 2» « © « » 390 " 
Tobacco Beetle . . : 2... 2 « 7 | Remedial Measures . . . - « « « 37 é 
Losses Due to the Tobacco Beetle. . . 7 | Preventive Measures ...-.-+ +. 53 3 
Distribution and Dissemination . . . 9 | Summary and Recommendations . . .« 68 ¥ 
Origin and History . ... . =. « « 10] Biblfography . . . . 2. 2 2 « «© « » 69 i 
WASHINGTON | oa 
_ GOVERNMENT PRINTING OFFICE a 
| 1919 eae 
