KUNGL. SV. VET. AKADEMIENS HANDLINGAR. BAND 5l. N:0O 7. 15 
have had rather a rough surface. It accordingly appears to show a certain resem- 
blance with L. Ottonis on this point, but, at any rate, the absence of such tubercles 
is of no importance. Finally, both of them have holes, but such are also found in 
other fossil plants from Scania. 
To sum up, it can be said that there are many circumstances which speak for, 
and none which speak against, there being a connection between ÅA. Zeilleri and L. 
Ottonis. As I have already pointed out, the arguments are not perhaps fully suffi- 
cient, and consequently the matter cannot be regarded as quite settled. But on the 
presumption that the fossils in question belong to the same plant, it will not be out 
of place to say a word or two on its systematical position. 
No seeds concerning which there exists any sort of evidence of their connection 
with Antholithus Zeilleri have been found; but it must, I think, be presumed that 
it constituted the male reproductive organ of a seed-plant. 
Although our knowledge of the male organs of the Pteridosperms is too limited 
for us to be able to make any statement as to their possible variation of shape and 
anatomy, they do not seem to differ essentially from the isosporangia of Marattia- 
ceous Ferns. As, however, it is not known whether the structures here described corre- 
sponded to segments of fertile fronds, such as in the Pteridosperms, or whether they 
themselves constituted differentiated microsporophylls grouped together in a kind of 
flower, and as the female organs are not known either, it is perhaps better not to 
count the hypothetical plant in question to this group. It is conceivable that in 
Mesozoic times the Pteridosperms were succeeded by a plant-group in which the male 
and the female sporophylls had reached a higher stage of evolution, and were more 
differentiated from the vegetative foliage still having filicinean habit. 
Antholithus Zeillert is only known from the Rhaetic deposits of Scania — to be 
more exact from the plant-bearing layer 4 (zone with Lepidopteris Ottonis). It occurs 
most frequently in the ”Blätterkohle” at Bosarp but also in solitary specimens at 
Billesholm and Bjuf. 
