36 CARL SKOTTSBERG, MARINE ÅLQM 1. PILEOPHYCEiE 



Scytosiphon (Ag.) Thur. 

 S. lomentaria (Lyngb.) J. Ag. 



Chiloé: Quemchi (St. 30, 19. 7. 08). W. Patagonia: Puerto Bueno, 

 sublit. on Mytilus, 1—3 m. (St. 27, 3. 6. 08, gt.)'; Puerto Riofrio (St. 28, 13. 6. 08.); 

 Isla Atalaya, lower litoral, common (St. 25, 25. 5. 08, gt.); Islas Evangelistas, in 

 pools, a very narrow form (St. 26, 26. 5. 08, gt.). Fuegia: Ushuaia, litoral, very 

 narrow fronds (St. 46, 14. 3. 09, gt.). Falkland Islands: Stanley Harbour, 

 litoral on pebbles (St. 1, 1. 11. 07, gt.) ; Cape Pembroke, common on Mytilus in tide- 

 pools (St. 3, 7. 1. 08, gt.); Port Louis (St. 11, 7. 2. 08, gt.); West Falkl., near Half- 

 way Cove (St. 4, 21. 11. 07). ■ - Groups of hairs, sometimes sunk below the surface, 

 resembling those figured in fig. 14 1 (Corycus) occur in various plants. I have not 

 seen any real cryptostomata, such as mentioned in Ant. Meeresalg. T, p. 34. 



Distribution: Aret. — Mediterr., Pacif. Ocean, Circump.-subantarct. 



Cladothele Hook. fil. et Harv. 



C. Decaisnei Hook. fil. et Harv. — Fig. 15 a — f. 



Falkland Islands: Cape Pembroke, in tide-pools, scarce (St. 3, 7. 11. 07, 

 sp. gt.) 



In Journ. of Botany XXIX Cladothele was reduced to Stictyosiphon by G. 

 Murray. He pointed out the conformity in anatomical strueture and referred to 

 Reinke^ excellent figures of the latter. In 1907, I was unable to form an indepen- 

 dent opinion on the subject, but with new and well preserved material at hand I 

 have arrived at the conclusion that the genus Cladothele must be restored. My specimens 

 are much more in accordance with the original figures of Hooker than with those 

 of Murray, which is the more remarkable as Murray regarded the former as less 

 accurate. The attention of the authors in »Flora Antarctica» was especially drawn 

 to the peculiar »papilla?» that cover the whole surface of Cladothele. Murray did 

 not get a good view of them and concluded that numerous empty gametangia had 

 suggested the idea of the papillae or »Codium-like cells». In my material these pa- 

 pillse, i. e. the strongly vaulted assimilatory cells, the development of which may be 

 understood from my figures, are very prominent, being free to V* or even 3 /* of their 

 length. This difference from Stictyosipho?i is, however, not sufficient to justify the 

 generic separation. But a study of the young apices revealed another difference. In 

 the latter, each branch terminates in a hair, the growth is trichothallic. In Clado- 

 thele every branch ends in an apical cell (fig. 15 a). If we follow Kjellman's prin- 

 ciples, as they were exposed in Nat. Pflanzenfam., this difference brings the two 

 plants not only to different genera, but to different orders. As the apical cell is 

 not an initial in the true sense, but becomes less active at an early date, Cladothele 

 might well find a place with the Dictyosiphonea? of Oltmanns (Morph. und Biol. 

 II, p. 367), as the resemblance to Delamarea is unmistakable. However, it deserves 

 to be mentioned that Kjellman described his Phlozospora (Stictyosiphon) pumila 



