12 HALLE, ON THE SWEDISH SPECIES OF SAGENOPTERIS PRESL AND ON HYDROPTERANGIUM NOV. GEN. 



hardly be doubted in regard to the great resemblance in essential characters. A 

 pedicel has, however, not yet been observed in these specimens. The number of 

 ridges and furrows is about the same as in the specimens from Bjuf, 12 — 14 (see 

 figs. 3, 4, pl. 2). They also radiate from the dorsal edge in the same manner. It 

 seems most probable, that these ridges correspond to the träns versly arranged pla- 

 centse in the sporocarps of Marsilia. Pl. 2, fig. 12 shows the impression of the out- 

 side of a capsule. The impression is covered by a carbonaceous film, the surface of 

 which probably represents the inner side of the capsule-wall. The ridges, appearing 

 on the carbonaceous lamina, would evidently have formed elevations on the inner 

 surface of the capsule-wall. Their number is somewhat greater than in the other 

 specimens. As seen in figs. 12—14 some of the ridges divide in the upper part. In 

 some places there is seen, between two ridges, a faint but fairly distinct vein, the 

 course of which can however be traced only a short distance. If the ridges really 

 represent - - as intimated above — the placenta? of a sporocarp, the arrangement of 

 veins and placentse would be more in accordance with the genus Regnellidium Lindm. 

 — considered by Prof. Lindman as a more primitive type — than with Marsilia. 

 The state of the material does, however, not permit of any sure conclusions in this 

 respect. A fact, not to be accounted for by a comparison with Marsilia or Regnelli- 

 dium, is the occurrence on the interiör casts of the Bjuf-capsules of narrow, projecting 

 ridges, which would correspond to furrows on the inner side of the wall. 



The specimen pl. 2, fig. 12 is of interest also in another respect. Pieces of the 

 carbonaceous lamina may be detached and, after suitable treatment, examined micro- 

 scopically. The lamina is seen to consist, in its present state, of two cuticularized 

 layers, separated by desintegrated remains of the interiör tissue. One of the two 

 cuticles (the outer one?) is better preserved than the other. The cells are polygonal 

 or rounded, with thick walls. The most striking feature is the occurrence of large 

 pores or stomata, which appear very clearly after staining the cuticle with saffranine 

 (pl. 3, figs. 12 — 15). The pores have about the same size as the epidermal cells and 

 are generally rounded or polygonal in shape. No distinct guard-cells are developed, 

 but the pore is surrounded by a ring of normally shaped epidermal cells. These 

 pores present an obvious resemblance to the »macrostomata» of the Marsilia-spoTo 

 carp. Some of the pores are more oval in shape and show some indistinct, projecting 

 remains of the cuticle round the borders. Such pores may possibly represent desinteg- 

 rated stomata of the common type, also occurring on the sporocarps of Marsilia 

 and Regnellidium. No typical stomata have, however, been observed with certainty. 



The occurrence of a cuticularized layer also on the inner side of the capsule- 

 wall is somewhat surprising and would be more in accordance with a f oliar nature 

 of the object. It is, however, not certain, that this layer represents a real epidermis, 

 as the cuticle - though of considerable thickness — is too desintegrated to give a 

 clear idea of its structure. It was, besides, impossible to decide, whether pores occur 

 also in the inner cuticle. 



Though the carbonized remains of the capsule have been examined microscop- 

 ically in several specimens, no spores have been observed which belong with cer- 



