FIONA. 



The anterior portions of the great hepatic channels are apparently connected with two 

 folliculated glandular bodies {g g), much and irregularly sacculated. These bodies are 

 united to the skin, one on each side, near the region of the stomach ; and probably form the 

 inner walls of those portions of the channels. Amidst the cellular tissue at the extremity of 

 the body, behind the ovary, there is likewise a glandular substance, (//), of a reddish colour, 

 folliculated and apparently ramified, in connexion with the branches of the hepatic canals 

 within the skin. These branches, at the posterior portion of the body, probably form a 

 sort of network of tubes across the dorsal aspect. Such, perhaps, may be inferred from the 

 appearance they present (/), when the skin of the back is divided down the median line. 



The arrangement of the hepatic canals differs from that which prevails in the Eolididce. 

 In Eolis, Embletonia, Doto, Dendronotus, Lomanotus, and Jnfiopa, the principal canals lie free 

 in the visceral cavity, and in all of them there is a median posterior trunk. In this genus 

 there is no such trunk ; and the canals are almost entirely within the skin. In this respect 

 Fiona resembles Hermcea, in which the whole of the hepatic ramifications are apparently 

 connected with the skin, and there are only two principal trunks, which pass down the 

 sides of the back. It is evident, however, that the digestive system alone sufficiently 

 distinguishes Fiona from all the above genera, not excepting Hermcea. 



The hepatic glands are large, nearly filling the papillae : they are slightly and irregularly 

 sacculated, with the inner surface of the investing membrane lined with a dark granular 

 substance; above, this substance is very abundant, forming a dense mass; below, the 

 membrane in some of the papillae is entirely devoid of it. We have failed to detect any ovate 

 vesicle like that of Eolis in the apex of the papillae, neither have we been able to determine 

 whether or not the apex is perforated. 



There are two large, much folliculated glands (fig. 8 c), lying beneath the stomach and 

 extending almost half way down the body. These terminate in front each in a slender duct, 

 (d, d), which sinks into the fleshy frontal margin of the foot. At first sight, these organs have 

 the appearance of being connected with the digestive system, and we originally looked upon 

 them as salivary glands ; but further experience induces us to believe that they are for the 

 purpose of secreting mucus, which probably passes out in front of the foot, lubricating the 

 surface over which the animal is moving.* 



The reproductive organs are placed further forward than usual, and have two external 

 orifices (fig. 1 a, b), one placed a little in advance of the other, on the right side of the head, 

 between and a little below the tentacles. The one in front is for the exsertion of the 

 untromittent organ, the other is rather small and is common to both the female and 

 androgynous apparatus. On laying open the dorsal skin, the reproductive organs are found, 

 as usual, to occupy much of the visceral cavity, having the stomach and intestine lying above, 

 and the buccal mass in front. They are formed on the type of those of Eolis. The mucus- 

 gland (fig. 4, n,n), is exactly of the same form, and the mode of union of the androgynous 

 parts with the oviduct and testis is the same as in that genus. The only modification of any 

 interest is in connexion with the testis (/). We know of no other species in the whole of 

 the Eolididce, in which it is furnished with a distinct vas deferens. In this respect Fiona 



* The so-named salivary glands of Doto fragilis may perhaps prove to be merely mucus-glands of 

 a similar nature. 



