22 



epidermal cells of Epilobiam and some cyperaceous plants and in the endo- 

 dermic cells of Andropogonese. 1 * 



The above stated construction -type is to be met with invariably in all 

 the examined species of genuine Bamlusa, for examples, B. vulgaris, 

 B. nana, B. stenoslachya, B. Oldhami and some undetermined species from 

 Formosa. We learn from the figures by Eoss 2) that B. arundinacea is 

 al so provided with an O-s heath. S c h w e n d e n e r 3 } has already enlisted Bam- 

 lusa among the plants provided with O-shaped endodermic cells. 



As far as our own observations are concerned/ the cortex of the roots 

 of the species of Arundinaria and Phijllostachys shows a type of construc- 

 tion quite different from the one described above. The figures 16 and 17 

 (PI. I.) may serve as illustrations. We notice here that hypodermal cells 

 have extraordinarily thickened outer walls, and thus replace the epidermis 

 in it? functions, the latter being destroyed very early and falling off. The 

 peripheral sclerenchyma-Iayer of varying thickness passes over gradually to the 

 cortical parenchyma. The outer layer of the cortical parenchyma is always 

 thicker than the inner, which is traversed by a number of radially arranged 

 air-lacuna. Again the endodermic cells manifest a quite different mode of 

 thickening ; the apposition of new material occuring only on the inner and 

 radial walls, resulting in the formation of the so-called C-sheath. Sch- 

 wendener 5) has also stated that Arundinaria possesses a C-sheath. In 

 Arundinaria quad ran gidaris, Phyllostachys Kumasasa, etc., the innermost 

 cell-layer of cortical parenchyma may be regarded as a " Verstiirkungsring " 

 in Schwendener's sense, its inner walls being much thickened and 

 ligniried. 



As to the structure of the axial cylinder it is here to be noted that 

 in the roots of Arundinaria-iyye the number of internal phloem-strands 

 amounts usually to one half that of the peripheral strands, while in those 

 of Bambusa-type both strands are almost equal in number. G) Besides, the 

 form of the internal solitary sieve tubes of the latter is very characteristic, 

 showing in cross-sections regular oval outlines. 



l)Klinge, Vergl. tint, iib. Gramineen- u. Cyperaceenwurzeln. (Abstract.) 



2) Ross, loc. cit. 



3) Sc h \v endener , Sckutziclieide u. ilire Verstiirkungen. Gesammelte Bot. Mittheilungen. 

 Bin, p. 128. 



4) K. Shibata, Beitr. z. Waclistumsgeschichte d. Bainbusgewiichse. Jour. Sc Coll. Imp. 

 Univ. Tokyo, Vol. XIII, pt. 3, p. 444, 495. 



5) S c h w e n d e n e r , loc. cit. p. 128. 

 Gj K. Shibata, loc. cit. p. 447. 



