Courtesy Duke Univ. Sch. For. Courtesy Duke Univ. Sch. For. 
Figure 167.—Gallery of /ps calligraphus Figure 168.—Gallery of the /ps 
on limb of a_ shortleaf pine. grandicollis, southern pine engraver, 
on limb of loblolly pine. Note egg 
niches and incompleted larval mines. 
The adult is brown to black, is from 3.5 to 4.5 mm long, and has four teeth on 
each side of the declivity. Egg galleries, from three to six, radiate away from a 
central nuptial chamber in the phloem, deeply scarring the sapwood. Larval tunnels 
extend a short distance in the inner bark and end in pupal cells. Adults remain under 
the bark for a short period before emerging. While there, they make irregular, 
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