30 MISC. PUBLICATION 4 7 7, U. S. DEPT. OF AGRICULTURE 



ovate to narrowly obovate, broadly rounded, glabrate except for the 

 usually strigose and fulvous-tomentose midrib; veins few, branching 

 and anastomosing; petioles short or moderately long; fruit small, 

 pedunculate. 



Range: Guatemala to Panama. 



Includes: Q. pacayana (type) and Q. copeyensis. 



The series Pacayanae is a rather remarkable endemic group, which, 

 in its small glabrate leaves with few veins, is quite clearly distinguished 

 from other rain-forest species. Its strigose midrib, usually somewhat 

 tomentose beneath, suggests a relationship to the series Adatae with 

 which it is coincident in range, but its thick leaves, not markedly 

 narrowed basally, deny such a relationship. 



11. Quercus pacayana sp. nov. 11 



Large tree. Twigs 1.5 to 3 mm. thick, glabrate, tan with numerous 

 prominent pale lenticels, becoming quite gray. Buds round-ovoid, 

 about 3 mm. long, glabrous, light brown, the stipules about 4 mm. 

 long, subulate, caducous or sometimes persistent for a while. Leaves 

 deciduous, about 4 to 8 cm. long, 2 to 3.5 cm. broad, rather thick, 

 drying yellow, elliptic to usually narrowly obovate, broadly rounded, 

 cordulate to usually rounded at base, subentire or crenately low- 

 toothed especially above the middle, margins minutely revolute, 

 upper surface dull, glabrous or the midrib sparingly strigose, lower 

 surface glabrous except the strigose and fulvous-tomentose midrib; 

 veins about 10 or 12 on each side, much branched and obviously 

 anastomosing near the margin, somewhat impressed above, very 

 prominent (including the reticulum) beneath; petioles 4 to 12 (usually 

 about 6) mm. long, glabrate, dark reddish brown. Catkins and 

 fruit not seen, but the fruit probably small and pedunculate. (See 

 pi. 30.) 



Range: Volcan de Pacaya, Guatemala (1,800 to 2,400 m.). 



This species and the following comprise the series Pacayanae whose 

 relationships (or lack of them) are discussed above. Q. pacayana is 

 closely related to Q. copeyensis from which it is distinguished by its 

 very prominent reticulation and decidedly revolute leaf margins. 



Specimens examined: 



GUATEMALA.' — Dept. Guatemala: Volcan de Pacaya, above Las Calderas, 

 November 30, 1938, Standley 58467 (F[2] [type as sheet no. 989364], USNA); 

 Standley 58474 (F). 



12. Quercus copeyensis nom. nov. 12 



Quercus costaricensis f. kuntzei Trel., Mem. Nat. Acad. Sci. 20: 

 146. pi. 288. 1924 (pro parte— as to type only). 



Large tree to 15 m. in height or taller. Twigs moderate (2 to 3 

 mm.), somewhat fluted, strikingly yellowish- white, glabrous. Buds 

 round-ovoid, about 3 mm. long, glabrous, brown, the stipules persis- 

 tent for a short time, about 7 mm. long, ligulate, glabrous. Leaves 



n Quercus pacayana sp. nov. — Arbor grandis, ramuli 1.5-3 mm. crassi glabrati, gemmae 3 mm. longae 

 rotundo-ovoideae glabrae, folia decidua 4-8 cm. longa 2-3.5 cm. lata coriacea elliptica vel anguste obovata 

 apice rotundata basi cordulata vel rotundata subintegra vel crenata glabrata, costa strigosa vel fulvo-tomen- 

 tosa excepta, venis utrinque 10-12 subtus prominentibus, petioli 4-12 mm. longi glabrati. 



i 2 Quercus copeyensis nom. nov.— Arbor grandis, ramuli 2-3 mm. crassi flavi glabri, gemmae 3 mm. longae 

 rotundo-ovoideae glabrae, folia decidua coriacea 4-8 cm. longa 2.5-4 cm. lata ovata vel obovata apice rotun- 

 data vel obtusa basi subcuneata vel cordulata integra glabra, costa subtus ad basim excepta, venis utrinque 

 6-7, petioli 4-7 mm. longi glabri, fructus annuus pedunculatus, cupula 10-12 mm. lata. 



