THE CENTRAL AMERICAN SPECIES OF QUERCUS 35 



sistently stellate-pub erulent or villous, the surface flat-bullate; veins 

 about 12 to 16 on each side, rather regularly spaced and parallel, 

 obscurely branching and anastomising, passing into the teeth, some- 

 what impressed above, very prominent (including the reticulum) 

 beneath; petioles 3 to 7 mm. long, glabrate or persistently tomentose. 

 Staminate catkins? Pistillate catkins 1 to 7 cm. long, 2- to 4-flowered 

 distally on the tomentose or glabrate peduncle. Fruit annual, usually 

 evidently peduncled; cups about 18 mm. in diameter, hemispheric, 

 the scales lanceolate, basally thickened, puberulent or tomentose; 

 acorns ovoid, about 18 mm. long, glabrous, one- third or one-half 

 included. (See pis. 38 and 39.) 



Range: Guatemala, Honduras, El Salvador, and Nicaragua. 



The variety sublanosa is very difficult to distinguish adequately 

 from the Central American representatives of the typical form. The 

 inconstancy of its characters and the great number of intermediate 

 specimens would make it very desirable to combine the two. Yet 

 the typically thin expanded leaves of the variety sublanosa with 

 regularly toothed margins, sparingly pubescent under surface, and 

 many regular veins are hard to reconcile with the very thick subentire 

 and densely tomentose leaves of typical Q. peduncularis. It is signifi- 

 cant that the latter form is concentrated in southern Mexico and 

 Guatemala, whereas the typical form of the variety sublanosa is 

 most common in Honduras and to the south. The greater number of 

 intermediates are encountered in Guatemala. This might indicate 

 one of the often publicized "hybrid swarms" that have never con- 

 clusively been demonstrated to occur in Quercus. 



The leaf form of the variety sublanosa is also very confusing in 

 distinguishing it from Q. purulhana. Only the characteristic short 

 dense pubescence of the lower leaf surface in the latter species can be 

 depended upon to make the distinction. 



Specimens examined: 



BRITISH HONDURAS.— El Cayo District: Mountain Pine Ridge, Feb- 

 ruary 22, 1931, Bartlett 11656 (NY) ; San Agustin, July-August 1936, Lundell 6626 

 (F, Mi, NY [2], US); Lundell 6729 (F, Mi, NY, US. USNA). 



GUATEMALA. — Dept. Huehuetenango: near Chalchitan, January 14, 1939, 

 Standley 62577 (F, USNA) ; mountains southwest of Malacatancito, January 14, 

 1939, Standley 62623 (F, USNA). Dept. Baja Verapaz: between Salama and 

 Rabinal, May 31, 1904, Cook and Doyle 280 (US [2]); Cook and Doyle 286 (US). 

 Dept. Sacatepequez: Cerro de la Cruz, above Antigua, January 22, 1939, 

 Standley 63822 (F, USNA); Standley 68881 and 63382 (F). Dept. Jalapa: 

 Volcan Jumay, north of Jalapa, December 1, 1939, Steyermark 82423 (F, USNA); 

 Cerro Alcaba, east of Jalapa, December 2, 1939, Steyermark 32608 (F, USNA). 

 Dept. Jutiapa: Los Llanitos, near San Jose Acatempa, December 21, 1938, 

 Standley 60581 and 60588 (F). 



EL SALVADOR.— Dept. Chalatenango: El Jute, 1928, Calderon 2U5 

 (F, US). 



HONDURAS.— Dept. Yoro: El Portillo Grande, August 1937, von Hagen 

 1038 (F, NY) ; Aguan River Valley, vicinity of Coyoles, June 30, 1938, Yuncker, 

 Koepper, and Wagner 8180 (DeP, F, 111 [type of Q. aguana], NY). Dept. 

 Comayagua: hills above Siguatepeque, near El Achote, July 26, 1936, Yuncker, 

 Dawson, and Youse 6186 (DeP, F, MBG, Mi, US [isotypes of Q. achoteana]) ; 

 about Siguatepeque, July 18, 1936, Yuncker, Dawson, and Youse 5961 (DeP, F, 

 MBG, Mi, NY, US [isotypes as of Q. achoteana var. sublanosa]); about Siguate- 

 peque, February 18, 1928, Standley 56184 (AA, F, US). Dept. Tegucigalpa: 

 Monte de la Flor, Tegucigalpa, December 1937, von Hagen 1206 and 1133 (F, NY). 



NICARAGUA.— Prov. Bluefields: San Rafael del Norte, March 25-26, 

 1917, Miller and Griscom 28 (US). 



