THE CENTRAL AMERICAN SPECIES OF QUERCUS 43 



Series Andinae Trel., Mem. Nat. Acad. Sci. 20: 141. 1924. 



Medium-sized or large trees; twigs thick, from loose-tomentose 

 glabrate with very prominent lenticels; buds ovoid to subfusiform, 

 from loose-tomentose becoming glabrate; stipules caducous; leaves 

 deciduous or subevergreen, moderately large, lanceolate to oblanceo- 

 late, often acuminate, entire or rarely coarsely toothed about the apex, 

 quickly or tardily glabrate beneath; petioles short or moderately long; 

 fruit annual, subsessile, medium-sized or rather large, the cup scales 

 rather loosely appressed, fulvous-pubescent. 



Range: The Colombian Andes and adjacent Panama. 



Includes: Q. humboldtii Bonpl., the type. 



20. Quercus humboldtii Bonpl. in Humboldt and Bonpland, PL 

 Aequinoct. 2: 155. pi. 130. 1809. 



Quercus tolimensis Humb. and Bonpl., PI. Aequinoct. 2: 153. pi. 



129. 1809. 

 Q. almaguerensis Humb. and Bonpl., PI. Aequinoct. 2: 157. pi. 



181. 1809. 

 Q. lindeni A. DC. in DC. Prodr. 16 2 : 32. 1864. 

 Q. humboldtii var. lehmanniana Hieron. ex Trel., Mem. Nat. 



Acad. Sci. 20: 142. 1924. 

 Erythrobalanus humboldtii Schwarz, Notizbl. Bot. Gart. Mus. 



Berlin-Dahlem 13: 496. 1937. 

 E. tolimensis Schwarz, Notizbl. Bot. Gart. Mus. Berlin-Dahlem 



13: 496. 1937. 

 E. lindeni Schwarz, Notizbl. Bot. Gart. Mus. Berlin-Dahlem 13: 



496. 1937. 

 E. duqueana Schwarz, Notizbl. Bot. Gart. Mus. Berlin-Dahlem 



13: 495. 1937. 



Medium-sized or large tree. Twigs 2 to 4 or rarely 7 mm. thick, 

 fluted or subterete, from loosely fulvous-tomentose quickly or tardily 

 becoming glabrate, the numerous light lenticels raised and very 

 prominent. Buds about 5 mm. long, ovoid or elongating and acute, 

 from loosely tomentose becoming glabrate and dull brown but the 

 scales ciliate; the ligulate stipules rather early caducous. Leaves 

 subevergreen or clearly evergreen, rather thin but hard and coriaceous, 

 10 to 20 cm. long, 3 to 7 cm. broad, lanceolate to elliptic or oblanceo- 

 late, acute or long-acuminate at apex, cuneate to rounded at base, 

 entire or rarely coarsely few-toothed near the apex, margins minutely 

 cartilaginous-re volute, flat or minutely crisped, upper surface glabrous 

 and somewhat shining or the base of the midrib persistently tomentose, 

 lower surface glabrate or rather persistently floccose especially along 

 the midrib, opaque or somewhat shining, not bullate; veins about 12 

 to 16 on each side, branching and very obviously anastomosing near 

 the margin, impressed above but slightly raised within the grooves, 

 very prominent beneath, the reticulum rather inconspicuously raised 

 beneath, less so above; petioles 4 to 10 or 15 mm. long, glabrate or 

 persistently tomentose. Staminate catkins 8 to 15 cm. long, rather 

 loosely flowered, the rachis sparsely villous, the obtuse anthers well 

 exserted from the villous perianth. Pistillate catkins about 1 cm. 

 long, 1- or 2-flowered. Fruit annual, solitary or paired on a peduncle 

 3 to 10 mm. long and 3 to 6 mm. thick with very prominent lenticels; 

 cups 2 to 3 cm. broad, deeply or shallowly cup-shaped, the margins 



