44 MISC. PUBLICATION 4 7 7, U. S. DEPT. OF AGRICULTURE 



inrolled or not, the scales narrowly ovate, obtuse, rather loosely 

 appressed, short -fuTvous-tomentose ; acorns 2 to 3 cm. long, 1.5 to 2 

 cm. broad, round or ovoid, from loosely silky-tomentose becoming 

 glabrate and brown, one-half or usually one-third included. (See pis. 

 54 to 57.) 



Range: Colombian Andes and adjacent Panama (1,500 to 3,400 

 m.); type from Popayan. Colombia (Bonpland 2083). 



Quercus h umboldtii is readily distinguished from other annual-fruited 

 species of the subgenus Erythrobalanus by its large lanceolate or 

 oblanceolate and usually entire leaves and its rather large cups with 

 fulvous-tomentose loosely appressed scales. It is related to Q. 

 costaricensis in the nature of its fruit and in bearing the aborted ovules 

 laterally or basally in the acorn, but the vegetative characters of the 

 two species are so utterly different as to indicate at most a rather 

 remote relationship. 



Until recent years there has not been available for study a very 

 complete series of Andean Quereas. The few collections at the dis- 

 posal of Humboldt and Bonpland and later of de Candolle were 

 insufficient to reveal their identities as one polymorphic species 

 differing only in the luxuriance of the foliage and persistence of 

 t omentum along the midrib beneath. A form with twigs and under 

 surface of leaves densely buff-tomentose and tardily glabrate {Perez 

 and Cuatrecasas 8100) is about representative of the plant named 

 Q. tolimensis except that the specimen here cited also has rather 

 small leaves. Tendencies toward this extreme pubescence occur 

 here and there in the range of the species. It is very doubtful that 

 this variation represents more than a casual form. Erythrobalanus 

 duqueana differs even less than the other proposed species, and to 

 recognize it as distinct would necessitate the recognition of numerous 

 additional minute variants as yet undescribed. 



A specimen in the Xew York Botanical Garden (Hayes 830) is 

 labeled merely " Panama. 1859-1860." The plant clearly belongs 

 to this species, but no other collections from Panama have ever been 

 found. Xo record of Hayes having collected in Colombia was 

 encountered. It is very likely that the Hayes specimen was taken 

 in lower Panama near Colombia, since no specimens of this species 

 have been turned up in the recent intensive explorations of the 

 Chiriqui region. 



Specimens examined : 



PANAMA. — Prov. Dariex (?) : without data. 1859-1860. Haues 830 (NY). 



COLOMBIA.— Without locality. 1783-1808, Mutis 5103 (F). Dept. San- 

 taxder: Eastern Cordillera, Mesa de los Santos, December 11-15, 1926. Killip 

 and Smith 15296 (NY, US): vicinity of Las Vegas, December 21-23, 1926, Killip 

 and Smith. 15895 (NY, US); Rio Surata valley, above Surata, January 5-6. 1927. 

 Killip and Smith 16744 (NY, US); west slope of Paramo de las Puentes, above la 

 Baja. January 25-31, 1927, Killip and Smith 18267 (NY, US); vicinity of Charta, 

 February 1-11, 1927, Killip and Smith 19078 (F, NY, US). Dept. Boyaca: 

 nearTunja, 1849, Linden 1325 (NY [2] [isotvpes of Q. lindeni]): Maripi, Julv 21, 

 1936, Garcia 4853 (US): Bosques de Arcabuco, February 24. 1940. Perez and 

 Cuatrecasas 8100 (US [2]). Dept. Cuxdixamarca: "El Colegio," February 

 1916, Dawe 55 (US); Fusagasuga, February 1876. Andre 171 (F, NY); Februarv 

 5, 1876, Andre 1433 (F) : Tequendama, October 28, 1917, Pennell 2640 (F, NY, 

 US) ; below falls of Tequendama on Bogota River, December 19, 1929, Xiemeyer 

 210 (US): Salto de Tequendama, October 1-3, 1938, Cuatrecasas 113 (US); 

 Caparrapi, June 8-13, 1939, Garcia 7 746 (US). Com. Caqueta: Cordillera 

 Oriental, eastern slope, Quebrada del Rio Hacha, Cajon de Pulido, March 26, 

 1940, Cuatrecasas 8784 (US); Sucre, April 4, 1940, Cuatrecasas 9076 (US). Dept. 



