54 MISC. PUBLICATION 4 7 7, U. S. DEPT. OF AGRICULTURE 



Standley 56398 (AA, F [type of Q. siguatepequeana]). Dept. Tegucigalpa: 

 Mont, de la Flor near Tegucigalpa, December 13, 1937, von Hagen 1220 (F, NY). 

 COSTA RICA.— Prov. San Jose: hills about Santa Maria, April 20, 1928, 

 Stork 1500 (F). Prov. Alajuela: Volcan Poas, March 31, 1907, Pittier 2036 

 (NY, US). 



27. Quercus eugeniaefolia Liebm., Overs. Danske Vidensk. Selsk. 

 Forhandl. 1854: 185. 1854. 



Quercus bumelioides Liebm., Overs. Danske Vidensk. Selsk. For- 

 handl. 1854: 188. 1854. 



Q. eugeniaefolia f. petiolata Trel., Mem. Nat. Acad. Sci. 20: 161. 

 pi. 316. 1924. 



Small or large tree, reaching 30 m. Twigs 1.5 to 2.5 mm. thick, 

 fluted, from sparsely loose-tomentose soon glabrate and glossy 

 reddish brown with rather prominent lenticels the second year. Buds 

 3 to 4 mm. long, fusiform, very acute, scales fulvous-ciliate about the 

 apex or glabrate, dull brown; the ligulate stipules early caducous. 

 Leaves subevergreen or distinctly evergreen, normally thick, in any 

 event quite hard and leathery, 6 to usually 9 or even 15 cm. long, 1.5 

 to 3.5 cm. broad, linear-lanceolate to narrowly subelliptic, apex 

 normally acuminate and aristate-tipped, sometimes long-acuminate or 

 merely acute or eveu individually rounded, base cuneate or attenu- 

 ately acute to rounded, entire, margins rather coarsely revolute (in 

 fully matured leaves) and usually finely crisped, upper surface glossy 

 and glabrous or minutely stellate-pubescent at the base of the midrib, 

 lower surface glabrous or the midrib sparsely stellate-pubescent toward 

 the base, the lamina dull and often waxy-glaucous in age; veins about 

 12 to 15 on each side and often with evanescent intermediates, issuing 

 at an angle of about 75 or 80 degrees, repeatedly branched and anasto- 

 mosing near the margin, impressed above but somewhat raised within 

 the grooves, more prominently raised beneath, the reticulum very 

 slightly raised on both surfaces; petioles 2 to 5 or even 7 mm. long, 

 rather distinctly winged, glabrate or sparsely stellate-pubescent for a 

 time. Staminate catkins about 4 cm. long, loosely flowered, sparsely 

 crisped- villous. Pistillate catkins? Fruit annual, small, solitary on 

 glabrous peduncles 3 to 8 mm. long; cups about 7 to 11 mm. broad, 

 about 5 mm. deep, cup-shaped or shallower, rounded or slightly con- 

 stricted at the base, scales ovate, broadly rounded, very closely ap- 

 pressed, sericeous-pub erulent except the glabrous brown apex and 

 margins; acorns 10 or 11 mm. long, 9 or 10 mm. broad (much smaller 

 and more slender with narrower cups in younger fruit which often 

 appears fully grown), ovoid to subrotund, fulvous-sericeous-puberu- 

 lent, light brown where abraded, about one-fifth or one-fourth in- 

 cluded; abortive ovules apical. (See pis. 71 and 72.) 



Range: Mountains of central Costa Rica to upper Panama (1,400 

 to 2,400 m.) ; type from Costa Rica {von Warscewicz C) without further 

 data, 



Trelease regarded Q. eugeniaefolia as related to Q. seemanni from 

 which it is often difficult to distinguish but to which it is not closely 

 related. It is distinguished from Q. seemanni by its prominently 

 revolute leaves with veins issuing at an angle of 75 or 80 degrees 

 (although this broad angle is rarely seen in very narrow leaves in 

 Q. seemanni) and its abortive ovules strictly apical. Q. eugeniaefolia 

 is probably most closely related to Q. sapotaefolia from which it is 



