THE CENTRAL AMERICAN SPECIES OF QUERCUS 57 



Quercus tonduzii is distinguished from Q. borucasana in the dis- 

 cussion under that species. From Q. eugeniaejolia, to which it is not 

 closely related but which it most confusingly resembles, it may be 

 separated by its veins which issue at an angle of about 45 degrees, its 

 larger acorns, and its rather loosely appressed cup scales. 



Neither Q. borucasana nor Q. tonduzii is clearly understood. The 

 former is known from three rather fragmentary collections and the 

 latter from a single but complete collection. This is not enough to 

 establish the amount of intraspecific variation or the dependability 

 of the characters in this rather difficult group of species. 



Specimens examined: 



COSTA RICA. — Prov. Alajuela: Volcan Poas, Achiote, November 1896, 

 Tonduz 10788 (US [2] [isotypes]). 



Series Rapurahuenses Trel., Mem. Nat. Acad. Sci. 20: 143. 1924. 



Series Citrijoliae Trel., Mem. Nat. Acad. Sci. 20: 147. 1924. 



Small or large trees; twigs slender, glabrate, buds 2 to 4 mm. long, 

 ovoid, glabrate; leaves evergreen or deciduous, rather small, lanceolate, 

 often long-acuminate, entire; petioles rather short; fruit annual, 

 rather small or medium-sized, cups cup-shaped or saucer-shaped, 

 the scales appressed, acorn usually covered at base only; abortive 

 ovules basal, lateral, or apical. 



Range: Costa Rica and upper Panama. 



Includes: Q. seemanni Liebm. (including Q. rapurahuensis Pittier, 

 the type of the series), and Q. gulielmi-treleasei C. H. Mull. 



This series is made up of moderate-sized, entire-leaved, annual- 

 fruited species with short petioles and acute or acuminate leaf apices. 

 It is characterized by appressed cup scales and rather small fruit. 

 Its relationship to other series is not clear except insofar as its entire 

 leaves and basal abortive ovules suggest the series Andinae and 

 Costaricenses , and from these its closely appressed cup scales clearly 

 distinguish it. The series Aristatae is more difficult to distin- 

 guish, but its distinctness is indicated by its even smaller leaves and 

 fruit and its cup scales exceedingly closely appressed and abortive 

 ovules apical. 



30. Quercus seemanni Liebm., Overs. Danske Vidensk. Selsk. Forhandl. 



1854: 188. 1854. 



Quercus citrifolia Liebm., Overs. Danske Vidensk. Selsk. Forhandl. 



1854: 187. 1854. 

 Q. salicijolia var. seemanni Wenzig, Jahrb. K. Bot. Gart. Berlin 3: 



207. 1884. 

 Q. rapurahuensis Pittier ex Seemen, Bui. Herb. Boissier 2. ser. 4: 



654. 1904 (nom. nud.): ex Trelease, Mem. Nat, Acad. Sci. 20: 



143. pL 275. 1924 (description). 

 Q. boquetensis Standi., Field Mus. Bot. Ser. 22: 13. 1940. 

 Q. chiriquiensis Trek, in Herb. 



Small or moderately large tree. Twigs 1 to 2.5 mm. thick, fluted, 

 from sparsely pubescent soon glabrate and dark reddish brown with 

 prominent light lenticels. Buds 2 to 4 mm. long, ovoid or elongating, 

 acute, glabrous, light brown; the ligulate stipules soon caducous. 

 Leaves subevergreen, thin but rather hard, 4 to usually 8 or even 16 

 cm. long, 1 to usually 3 or 4 cm. broad, lanceolate or occasionally 



