THE CENTRAL AMERICAN SPECIES OF QUERCUS 21 



Series Oocarpae Trel., Mem. Nat. Acad. Sci. 20: 43. 1924. 



Large trees with rather coarse villous or glabrate twigs; buds ovoid 

 or subglobose, the stipules caducous or persistent; leaves large, 

 evergreen(?), typically oblanceolate, coarsely toothed; fruit large, 

 cotyledons unequal. 



Range: Guatemala to Panama. 



Includes: Q. oocarpa Liebm. (type) and Q. pilarius Trel. 



4. Quercus oocarpa Liebm., Overs. Danske Vidensk. Selsk. For- 

 handl. 1854: 184. 1854. 



Q. warscewiczii Liebm., Overs. Danske Vidensk. Selsk. Forhandl. 



1854: 187. 1854. 

 Q. yunckeri Trel. in Yuncker, Field Mus. Bot. Ser. 17: 358. 1938. 



Small or large tree (6 to 25 m.). Twigs rather coarse (3 to 5 mm.), 

 fluted, from densely fulvous-tomentose becoming glabrate and gray or 

 light brown with a few scarcely evident lenticels. Buds oblong, 

 acute, about 4 mm. long, glabrous, with ligulate villous stipules 10 to 



13 mm. long and persistent. Leaves apparently evergreen, thin and 

 papery or rather firm, 10 to 25 or rarely 30 cm. long, 3 to 10 or even 



14 cm. broad, oblanceolate to obovate or narrowly elliptic, attenuately 

 acute at apex, narrowly rounded or cuneate at base, undulately or 

 sharply low-dentate except the entire base, margins very minutely 

 somewhat revolute, upper surface somewhat shining, glabrous or 

 usually glabrescent from minutely puberulent, the midrib often 

 persistently fulvous-tomentose, lower surface dull, persistently and 

 sparsely villous especially on the veins and reticulum ; veins about 14 

 to 22 on each side, much branched and obviously anastomosing near 

 the margin but eventually passing into the teeth where those are 

 present, slightly impressed above and very prominent (including the 

 reticulum) below; petioles very short (3 to 5 mm. long), densely or 

 sparsely fulvous-tomentose. Catkins? Fruit annual, rather large, 

 solitary or paired on a peduncle 5 to 18 mm. long and 5 to 7 mm. 

 thick; cups 3 to 4 cm. broad, openly goblet-shaped or cup-shaped, 

 the scales triangular-ovate to oblong and narrowly obtuse, densely 

 short-pubescent, loosely appressed; acorns depressed-subglobose or 

 elliptic, 2.5 cm. long and 3 cm. broad or (in the Panama material) 

 4 cm. long and 2.5 cm. broad, one-half or only one-fourth included. 

 (See pis. 10 and 11.) 



Range: Guatemala, Costa Rica, and upper Panama (1,400 to 1,900 

 m.) ; type from Guatemala (von Warscewicz 50a, without further- 

 locality) . 



Quercus oocarpa is the type species of the series Oocarpae which 

 is not closely related to the other large-fruited series and is character- 

 ized by its oblanceolate, toothed, attenuately acute leaves with 

 markedly short and inconspicuous petioles. The thin leaves, villous 

 beneath, distinguish Q. oocarpa from Q. pilarius. 



Specimens examined: 



GUATEMALA. — Deft. Alta Verapaz: near Finca Sepacuite, April 10, 1902, 

 Cook and Griggs 572 (US). Dept. Baja Verapaz: Valley of Santa Rosa, June 5, 

 1904, Cook 220 (US). Dept. Guatemala: Volcan de Pacaya, between San 

 Francisco Sales and base of the active cone, December 20, 1940, Standley 80624 

 (USNA). Dept. Chiquimula: Volcan Quezaltepeque, northeast of Quezalte- 

 peque, November 8, 1939, Steyermark 31%85 (F); Volcan Ipala, near Amatillo, 

 October 25, 1939, Steyermark 30525 (F); upper slopes of Montana Tajuran, 

 vicinity of El Barriol, October 28, 1939, Steyermark 30807 (F). 



