66 MISC. PUBLICATION 4 7 7, IT. S. DEPT. OF AGRICULTURE 



Large tree. Twigs 1 to 2 mm. thick, fluted, glabrous or from sparsely 

 stellate-pubescent glabrate and reddish brown with rather inconspicu- 

 ous lenticels. Buds about 3 or rarely 4 mm. long, scarcely 2 mm. 

 broad, ovoid, acute, sparsely pubescent or glabrate, light brown; the 

 ligulate stipules early caducous, 5 to 7 mm. long, sparsely pubescent 

 dorsally. Leaves evergreen, rather thick and hard or rarely thin, 3 or 

 usually 5 to 10 or even 15 cm. long, 1 or usually 2 to 3 or even 4.5 cm. 

 broad, typically lanceolate to elliptic or narrowly ovate, acute or acu- 

 minate, nor markedly aristate-tipped, cimeate to rarely attenuately 

 narrowed at base or in short-petiolate juvenile forms or stump-sprouts 

 rounded or cordulate, entire or obscurely few-toothed toward the apex, 

 juvenile forms coarsely serrate, margins minutely cartilaginous-revo- 

 lute and somewhat crisped, upper surface somewhat shining, glabrous 

 or slightly pubescent along the midrib, lower surface less shiny or 

 opaque, glabrous or with rather prominent domatia in the axils of the 

 principal veins, the lamina quite smooth (not bullate) ; veins about 10 

 to 14 on each side, branching and rather obviously anastomosing near 

 the margin, slightly raised above and more prominent beneath, the 

 reticulum slightly raised on both surfaces; petioles 5 to 10 or 20 mm. 

 long, in juvenile forms often scarcely 2 mm. long, usually rather prom- 

 inently winged. Staminate catkins about 3 cm. long, fulvous- or 

 gray-villous, loosely flowered, the oblong anthers well exserted. 

 Pistillate catkins 5 to 10 or even 20 mm. long, the glabrous peduncle 

 usually 2-flowered. Fruit biennial, solitary or paired on a peduncle 

 5 to 20 mm. long, 2 to usually 4 mm. thick, with large conspicuous 

 lenticels; cups 10 to 20 mm. broad, 7 to 12 mm. high, cup-shaped to 

 turbinate and constricted basally, the scales triangular to lanceolate or 

 ovate, the apices usually narrowed but rounded, sericeous-canescent 

 or locally glabrate; acorns 10 to 17 mm. long, 8 to 14 mm. broad elliptic 

 to ovoid, sparsely puberulent or glabrate, light brown, one-half or 

 only one-third included. (See pis. 91 to 93.) 



Range: Mountains of Guatemala where it is very abundant, the 

 Mexican State Chiapas, and perhaps also to be expected in Honduras; 

 type from Guatemala (von Warscewicz 1+7 and ^8) without further 

 data. 



Quercus acatenangensis is a highly variable species and therefore very 

 difficult to characterize. The degree of variability, however, is not 

 greater than that of other well-known species in which the segregation 

 of new species has proved unsuccessful (e. g., Q. macrocarpa Michx. 

 and Q. nigra L.). Q. donnell-smithii and Q. ambivenulosa differ in no 

 fundamental characters. The broad-leaved form on which the latter 

 is based merges directly into the elongate narrow-leaved form of typical 

 Q. acatenangensis. The specimen cited as the type of Q. donnell- 

 smithii (Smith 1967) is actually Q. sapotaejolia, but Trelease's concept 

 of Q. donnell-smithii is obviously based upon a specimen of Q. acatenan- 

 gensis. His illustration (pi. 318), although labeled in the explanation 

 as Smith 1967 and described as two variants of the type, is clearly 

 made from Smith 2629. Two sheets of this latter number are to be 

 seen in the United States National Museum and the New York 

 Botanical Garden, both quite clearly exhibiting the two variants il- 

 lustrated although neither sheet is the actual one photographed. The 

 description of the species is obviously based upon the illustrated but 

 mislabeled Smith 2629 rather than the cited number 1967. 



