6 



gelen danner Bunden, Fig. 15, e. Tab. II, Fig. 2, a. Denne 

 Rende strækker sig opad fra Svælgfladens nederste Parti til 

 lidt over Trefjerdedelen af samme og er bredest nedad, Tab. 

 II, Fig. 2, a. Den adskiller sig fra den øvrige Del af 

 Svælgvæggen væsentlig ved sin særegne Epithelbeklæd- 

 ning, der dannes af meget lange, næsten traadformige 

 Pidskeceller (Geisselcellen), som bave en yderst tynd Mem- 

 bran, ere 0.060'™' lange, 0.020'»» brede, og en aflang Kjerne., 

 0.004""» lang, 0.002'" m bred, der er placeret snart nedimod 

 den nederste Ende, snart paa Midten og snart bøiere oppe, 

 Fig. 27. Celleindboldet er meget klart, og idet hver Celles 

 fri Ende, der synes at være rig paa Protoplasmaindbold, 

 støder tæt til Nabocellens, fremkommer en skarp Rand, der 

 bar Udseende af en Cuticula, Tab. II, Fig. 2 b, hvorfra 

 udgaa en Række lange, ved Grunden tykke Flimmerbaar 

 (Ciher), som rage langt ind i Svælghulheden, Fig. 28. Tab. Il, 

 Fig. 2; enhver Celle bærer kun et saadant Haar, der er 

 0.040™» langt, Fig. 29. 



Ved Tversnit af Svælget paa Alcoholpræparater faar 

 man et eiendommeligt Billede af denne Svælgrendes Epi- 

 thel, — det ser ud, som om det bestaar af mange Lag 

 næsten elliptiske Celler, hvoraf de inderste, nærmest Hul- 

 heden, antage Cylinderformen og bære paa deres frie Ender 

 Elimmerhaar, Tab. II, Fig. 2 c. Kun ved Maceration 

 fremkom disse Cellers sande Form 1 . 



Udenibr Renden og saa langt, den naar op, er Svæl- 

 get beklæclt med Cylinderepithel, forsynet med sædvanlige 

 Cilier; men ovenfor Svælgrenden dannes Epithelbeklædningen 

 af Polypens Ectodermceller, der dog langs Rygsiden stræk- 

 ker sig langt nedover Svælget. Imellem Svælgets Cylinder- 

 epithel findes en stor Mængde kolbeformige Legemer, der 

 ligge dels spredte, dels i regelmæssige Rækker omkring 

 Længdefolderne, sjeldnere i Grupper, Tab. II, Fig. 2, d. 



De ere fra 0.020— 0.040'»'» lange og fra 0.010— 0.015"™ 

 brede, som oftest vandklare og se ud som Vacuoler, have 

 en langstrakt Hals med en Aabning paa Enden, Fig. 30. 

 Tab. II, Fig. 2, e. Ved Farvning fremkommer i den 

 nedre, brede Del en næsten rund Kjerne, 0.004""" i Gjen- 

 nemsnit, med Kjernelegeme, omgiven af en seig Masse, der 

 hyppig var trykket ud igjennem den før omtalte fine Aab- 



1 Dr. Blochman fra Heidelberg, eler i længere Tid har arbeidet 

 i Bergens Museum, viste mig Præparater fra den indre Tarmvæg 

 hos Brachiopoderne, hvor det samme Forhold fandt Sted. Af hans 

 Maqerationspræparater fremgile det tydelig nok, at Epithelbeklæd- 

 ningen bestod af lange, traadformige Celler, forsynede med lange 

 Cilier, imedens den paa Alcoholpræparater viste sig som ovenfor om- 

 talt, nemlig ligesom bestaaende af flere Cellelag. 



the broad part of the triangle forms the bottom, (Pl. I, fig. 15, e, 

 Pl. II, fig. 2, a). This channel extends itself upwards, from 

 the lowest part of the surface of the gullet, till a little 

 above the three-fourths part of it, and it is broadest below, 

 (PI. 2, fig. 2 a). It is distinguished from the rest of the 

 wall of the gullet principally, by its peculiar epithelial 

 covering, formed of very long, flagellated, almost filiform, 

 cells (geissel-cells) having an extremely thin membrane, 

 and measuring 0.060""» in length, and 0.020""» in breadth, 

 and containing an oblong' nucleus measuring 0,004'»'" in 

 length and 0.002""" in breadth, placed sometimes, down 

 towards the lowest extremity, sometimes, in the middle and, 

 sometimes, higher up, (PI. I, fig. 27). The cellular sub- 

 stance is very translucent and, as each cell's free extremity, 

 which appears to be rich in protoplasmic substance, ap- 

 proaches close to that of the neighbouring cell, a distinctly- 

 marked margin is produced, which has the appearance of a 

 cuticulum, (PL, II fig. 2, b), from which a series of long cilia, 

 thick at the root, proceed, extending far into the gullet- 

 cavity, (PL I, fig. 28, PL II, fig. 2). Each cell carries 

 only one such cilium measuring 0.040 7 "™ in length, (PL I, 

 fig. 29). 



On making a transverse section of the gullet, in pre- 

 parations preserved in alcohol, we obtain a characteristic 

 representation of this gullet-grooves epithelium; it appears, 

 as if it consists of many layers of almost elliptical cells, of 

 which the innermost, next to the cavity, take the cylinder- 

 form, and carry cilia on their free extremities, (PL 11, fig. 2. c). 

 Only after, maceration, did the true form of these cells 

 appear L 



Outside the channel, and as far up as it reaches, the 

 gullet is clad with cylinder-epithelium furnished with the 

 usual cilia, but above the. gullet- channel the epithelial 

 covering is formed by the ectoderm-cells of the polyps, 

 which extend however along the dorsal side far down the 

 gullet. In the gullets cylinder-epithelium, a great multitude 

 of clavi-form corpuscles are found, which are placed, 

 partly dispersed and, partly, in regular series about the longi- 

 tudinal folds, more rarely in groups, (PL II, fig. 2 if). 



They measure, from 0.020 — 0.040'"™ in length, and 

 from 0.010 — 0.015""" in breadth, and are most frequently 

 translucent, appearing like vacuoli; they have an elongated 

 neck with an aperture on the extremity, (PL I, fig. 30, 

 PL II, fig. 2, e). On staining, an almost spherical nucleus 

 appears in the lower, broad part; it measures 0.004""" in 

 diameter, and has its nucleus body surrounded by a tough 

 substance, which was frequently exuded from the minute 

 aperture previously spoken of. These corpuscles are not 



1 Dr. Blochman, from Heidelberg, who studied for a consider- 

 able time in Bergens Museum, showed me preparations from the 

 inner intestinal wall of the Brachiopods where the same relations 

 existed. From his macerated preparations it appeared, distinctly 

 enough, that the epithelial covering consisted of long filamentous cells 

 furnished with long cilia, whilst in preparations preserved in alcohol 

 it appeared as above described, viz. as if consisting of several cell- 

 ular layers. 



