746 MISC. PUBLICATION 42 3, U. S. DEPT. OF AGRICULTURE 



1. Lappula texana (Scheele) Britton, Torrey Bot. Club Mem. 5: 273. 



1894. 



Echinospermum texanum Scheele, Linnaea 25: 260. 1853. 

 Echinospermum redowskii (Hornem.) Lehm. var. cupulatum 



A. Gray, Bot. Calif. 1: 530. 1876. 

 Lappula heterosperma Greene, Pittonia 4: 95. 1899. 



Apache County to eastern Mohave County, 5,000 to 6,500 feet, 

 type of L. heterosperma from Peach Springs (Greene in 1889). Kansas 

 to Idaho, south to Texas, New Mexico, and northern Arizona. 



The range in Arizona, as given above, is that of typical L. texana, 

 with heteromorphic nutlets. The var. coronata (Greene) Nels. and 

 Macbr. (L. coronata Greene), with nutlets all alike in form, is found, 

 mostly at lower elevations, in Graham, Cochise, and Pima Counties, 

 type from mesas near Tucson (Pringle in 1884). A very similar form 

 is found in the northern Rocky Mountain States. 



2. Lappula echinata Gilib., Fl. Lithuan. 1: 25. 1781. 



Schultz Pass, Coconino County (Whiting 1173B). Widely dis- 

 tributed in the northern United States and Canada; introduced from 

 Eurasia. 



3. Lappula redowskii (Hornem.) Greene, Pittonia 2: 182. 1891. 



Myosotis redowskii Hornem., Hort. Hafn. 1: 174. 1813. 

 Lappula occidentalis (S. Wats.) Greene, Pittonia 4: 97. 1899. 

 Lappula leucotricha Rydb., Torrey Bot. Club Bui. 36: 676. 

 1909. 



Widely distributed in Arizona in the creosotebush, juniper, and pine 

 belts, 1,200 to 8,500 feet, usually in sunny places in disturbed soil, type 

 of L. leucotricha from near Tucson (Tourney in 1894) . Western United 

 States; Argentina; Asia. 



The species has many forms, but these do not seem to be correlated 

 with geography and more than 1 of them may occur in a given locality. 

 The typical form has the marginal appendages distinct or nearly so on 

 all 4 nutlets. This is less common in Arizona than var. desertorum 

 (Greene) Johnston (L. desertorum Greene, L. leucotricha Rydb.), in 

 which the appendages are more or less evidently confluent on 1 or 

 more of the nutlets in each fruit. 



6. HACKELIA 23 



Coarse perennial or biennial herbs; corolla white or blue, in naked 

 racemes; nutlets attached by a submedial areola to a pyramidal 

 gynobase; plants of meadows, thickets, and pine forests. 



Key to the species 



1. Corolla white or at most bluish only about the center, 5 to 10 mm. wide; 



marginal appendages of the nutlets commonly much united. 1. H. ursina. 



1. Corolla normally blue, 3 to 7 mm. wide; marginal appendages of the nutlets 



free or united only at base (2) . 



2. Middle stem leaves tending to be petiolate; branches usually few, elongating, 



spreading, only very rarely aggregated into a conspicuously cylindric 



inflorescence; plant slender 2. H. pinetorum. 



2. Middle stem leaves tending to be sessile; branchlets bearing mature cymes 

 numerous, short, together forming a leafy-bracted elongate terminal 

 panicle; plant coarse, rather strict 3. H. floribunda. 



23 Reference: Johnston, I. M. restoration of the genus hackelia. Gray Herbarium Contrib. 

 68: 43-48, 1923, 



