NORTH AMERICAN BOSTRICHIDAE 19 



reddish yellow, each elytron with three subrounded greenish-black 

 spots, and Lesne (1938) placed it as an aberration of quadrisignata. 



The markings on the elytra of this species are quite variable and 

 do not seem even to represent stable aberrations or varieties. In a 

 large series of specimens collected in Los Angeles County by Coquillett 

 on the same date, all four named forms are represented (maculata, 

 quadrisignata, cleroides, and sexguttata) . 



It is very difficult to give any distinct characters for separating 

 metadata and quadrisignata on account of the variable elytral mark- 

 ings. In maculata the elytral markings have a tendency to become 

 vittate, whereas in quadrisignata the dark ground color gradually be- 

 comes smaller, leaving the elytra simply reddish yellow and more or 

 less spotted with blue. In quadrisignata the thorax is more nearly 

 ovate and more shining and convex than in maculata. resembling 

 some species of Polycaon, whereas in maculata it is more depressed and 

 resembles some of the Cleridae. 



Subfamily ENDECATOMINAE 



Endecatomini LeConte, 1861, Smithsn. Inst. Misc. Collect. 3 (1): 207; Horn, 

 1878, Amer. Phil. Soc. Proc. 17 : 540 ; LeConte and Horn, 1883, Smithsn. Inst. 

 Misc. Collect. 507 : 227. 



Hendecatomklae Kiesen wetter, 1877, in Erichson, Naturgesch. Insect. Deut, 

 Coleopt., v. 5, pt. 1, pp. 7, 23-25; Lesne, 1921, Assoc. Franc, pour l'Avanc. des 

 Sci., Cong, de Strasbourg (1920) , p. 287 ; 1934, Soc. Ent. de France Bui. 39 : 174 ; 

 1935. Paris Mus. d'Hist. Nat. Arch. (vol. du Tricent.) (ser. 6) 12: 432-433. 



Hendecatominae Lesne, 1938, in Junk (pub.), Coleopt. Cat., pt. 161, pp. 4-5. 



This subfamily is represented by the genus Endecatomus Mellie. 

 Most early writers associated this subfamily or tribe with the Cisidae, 

 but Jacquelin-Duval (1859-1863) placed it in the family Lyctidae. 

 LeConte (1861) erected the tribe Endecatomini for Endecatomus 

 rugosus (Randall) and placed it in the family Bostrichidae, stating 

 that this tribe is a connecting link between the Anobiidae and 

 Bostrichidae. Lesne (1921, 1934, 1935) placed it as a subfamily of 

 the Bostrichidae. Forbes 4 stated that according to the wing folding 

 pattern it is a normal bostrichid and does not show any connections 

 with the Cisidae. Lesne (1935) stated that according to the sensory 

 pits on the antennae, buccal pieces, wings, tarsi, abdominal stigmae, 

 and male genitalia, Endecatomus is very distinctly connected to the 

 Bostrichidae. 



Genus ENDECATOMUS Mellie 



Endecatomus Mellie, 1847, Rev. Zool. 10: 108-109; 1848, Soc. Ent. de France 

 Ann. (ser. 2) 6: 206-209, 213-218, pi. 9, figs. 1-8; Redtenbacher, 1858, Fauna 

 Austriaca, Kafer, ed. 2, pp. CI, 571; 1874, ed. 3, pt. 2, pp. 67-68. 



Hendecatomus Bach, 1852, Kaferf. v. 2, pt. 3, pp. 100, 108; Schaum, 1S52, Cat. 

 Coleopt. Europae, ed. 4, p. 55 ; Dohrn, 1855, Cat. Coleopt. Europae, ed. 5, p. 56 ; 

 1856, ed. 6, p. 53 ; Lacordaire, 1857, Genera des Coleopt., v. 4, pp. 547, 549 ; 

 Jacquelin-Duval, 1859-1863; Genera Goleopt. Europe, v. 3, p. 235; Gemminger 

 and Harold. 1869, Cat. Coleopt.. v. 6, p. 1795; Stein and Weise, 1877. Cat. 

 Coleopt. Europae, p. 110; Kiesenwetter, 1877, in Erichson, Naturgesch. Insect. 

 Deut., Coleopt.. v. 5. pt. 1. pp. 23-25, tig. ; Horn, 187S, Amer. Phil. Soc. Proc. 17 : 

 540-541; Heyden, Reitter, and Weise, 1883, Cat. Coleopt. Europae. p. 125; 

 LeConte and Horn, 1883, Smithsn. Inst. Misc. Collect. 507: 227; Marchal, 1888, 

 Feuille Jeunes Nat. 18 (208) : 50; Seidlitz, 1889, Fauna Baltica, Kiifer. ed. 2. 



4 Forbes, W. T. M. wing folding patterns of coleoptera. N. Y. Ent. Soc. 

 Jour. 34: 99 (fig. 75). 1926. 



