Electron niicrojiraph shoMs foot-and- 

 niouth disease virus particles 

 (about one-millionth inch in diam- 

 eter). One polystyrene reference 

 particle is at left center. 



► 



► 



► 





1 



Pioneered in the use 

 study of animal viri 



Adapted vesicular slia 

 culture. 



Isolated vesicular stifcti 

 man being. 



Developed a tissue jti 

 mouth disease virus v\ 



Developed a line of 111 

 growth of the seven l|s 



Viewed foot-and-moi. 

 croscope, and descr; 

 23 millimicrons (ab 

 made up of a prol 

 ribonucleic acid (Rl 



Confirmed that the 

 the RNA core of F? 

 infectious portion. 



Established that ri 

 widespread in anima 

 does not affect inta 

 inactivate RNase. 



Showed that FMDV 

 vated by heating at i 



Demonstrated that R 

 may be obtained fro 



Found that FMDV i 

 infection in several 

 naturally susceptible 



V 



