anda 1957 (Chis.); Schwerdtfeger 1953 (Guate- 
909 (Chih.); Sociedad Botanica de México 1960 
‘ley and Steyermark 1958 (Guatemala); Zobel 
CNEWEE 
lished—P. H. Allen 1962 (EI Salvador); B. Hall- 
{ (Oax.); Mexico, Instituto Nacional de Investiga- 
restales 1964 (Gro.); E. Larsen 1962; R. McVaugh 
Lo). 
i michoacana Martinez MAP 50 
Michoacan pine 
Pinus michoacana Martinez, Mex. Inst. Biol. An. 
libel tro 4s Oa 
This pine grows at relatively low elevations in central 
and southern Mexico, from southern Durango and San Luis 
Potosi southeast to Chiapas. It is sometimes associated with 
its close relative, Pinus montezumae. 
\dditional sources: 
Published—Guzman and Vela 1960 
(Mich.) . 
Unpublished—L. W. Bryan 1964  (Oax.); B. Hallberg 
1964 (Oax., Chis.); E. Larsen 1962 (Mich., Dgo.) ; N. Sanchez 
Mejorada to L. W. Bryan 1963 (Mich.). 
(Zac.) ; Loock 1950 
Pinus pseudostrobus Lindl. MAP 52 
Pinus pseudostrobus Lindl., Bot. Reg. v. 25, 
Misc. 63. 1839. 
Pinus pseudostrobus, including the doubtfully distinct 
P. tenuifolia Benth. (not Salisb.) and P. oaxacana Mirovy, is 
widespread at moderately high elevations from Sinaloa and 
Nuevo Leon through central and southern Mexico, and in 
Central America as far south as northwestern Nicaragua. This 
variable species complex extends farther south than any other 
western yellow pine. In the western part of its range it over- 
laps the closely related P. douglasiana. 
Additional sources: 
Published—Aguilar 1961 (Guatemala); Allen 1955 (Hon- 
duras); Castanos 1962 (Oax.); Denevan 1961 (Nicaragua) ; 
Gentry 1946 (Sin.); Goldman 1951 (Mich.); Leavenworth 
1946 (Mich.); Loock 1950 (Mich.); Miranda 1961 (Chis.); 
Miranda and Sharp 1950 (Pue.); Pfeifer 1960 (Honduras) ; 
Shaw 1909; Sociedad Botanica de México 1960 (Ver.); Vogel 
1954 (Honduras) ; Wicht 1949 (Mich.); Zobel and Cech 1957 
(NES 
Unpublished—P. H. Allen 1962 (El Salvador); B. Hall- 
berg 1964 (Oax.); N. Sanchez Mejorada to L. W. Bryan 1963 
(Mich.) . 
Pinus douglasiana Martinez MAP 532 
Douglas pine 
Pinus douglasiana Martinez, Madrono 7: 4, t.l. 1943. 
This rather uncommon pine is widely scattered at middle 
elevations in western Mexico from extreme southern Sonora 
to Michoacan and México. Its range slightly overlaps that of 
its closest relative, the highly variable Pinus pseudostrobus. 
Additional sources: 
Unpublished—B. Hallberg 1964 (Oax.) ; 
1964 (Dgo.). 
Cc. E. Blanco 
Pinus teocote Schiede & Deppe MAP 33 
Pinus teocote Schiede & Deppe in Schlecht. & Cham., 
Linnaea 5: 76. 1830. 
Pinus teocote, including the weakly segregated P. herrerai 
Martinez, is one of the most widely distributed Mexican 
pines, ranging from Chihuahua and Coahuila south to 
Chiapas, and just entering western Guatemala. In southern 
Mexico it overlaps P. lawsonii, a related species. 
18 
Additional sources: 
Published—Aguilar 1961 (Guatemala); Matuda 1950 
(Chis.) ; J. Rzedowski 1956 (S.L.P.); G. C. Rzedowski 1960 
(S.E-P.);) Shaw) 1909 (SE: P);" Wicht7 1949) 5 ((Michs)p- rele VValle 
liams 1939 (Oax.). 
Thesis—James H. Maysilles 1959. Floral relationships 
of the pine forests of western Durango, Mexico, Ph.D. thesis, 
Univ. Michigan, 165 pp., illus. 
Unpublished—C. E. Blanco 1964 (Coah.); B. Hallberg 
1964 (Oax.); F. Medellin-Leal 1964 (S.L.P.); O. Zarzosa EL. 
1964 (Dgo.). 
Pinus lawsonii Roezl MAP 53 
Lawson pine 
Pinus lawson Roezl ex Gord., Pinet. Sup. 64. 1862. 
This pine is confined to southern Mexico from Jalisco 
to Veracruz and south to Oaxaca. Its range overlaps that 
of the related Pinus teocote. 
Additional sources: 
Published—Loock 1950 (Mich.) . 
Unpublished—B. Hallberg 1964 (Oax.); N. Sanchez 
Mejorada to L. W. Bryan 1963 (Mich.) . 
SUBSECT. SABINIANAE Loud. 
Pinus sabiniana Dougl. MAP 54 
Digger pine 
Pinus sabiniana Dougl. ex D. Don in Lamb., Descr. 
Genus Pinus. Ed. 3 (8°), v. 2, unnumbered p. 
between p. 144 and p. 145, t. 80. 1832; 
Dougl., Linn. Soc. London Trans. 16: 749. 1833. 
Digger pine is confined to California. It 1s characteristic 
of the dry foothills of the Sierra Nevada and the Coast Range. 
Sources: 
Published—Griffin 1964. 
Unpublhished—]. R. Griffin 1964. 
Pinus coulteri D. Don MAP 55 
Coulter pine 
Pinus coultert D. Don, Linn. Soc. London Trans. 
17: 440. 1836. 
Coulter pine is restricted to the coastal mountains of 
central and southern California and northern Baja California. 
It overlaps two related species, Pinus jeffreyi and P. sabiniana, 
and is known to hybridize in nature with P. jeffrey: (Zobel 
1951). The distribution of the species in California is based 
entirely on Vegetation-Type survey information. ‘The sources 
listed below all pertain to Baja California. 
Additional sources: 
Published—Goldman 1916; Martinez 1948. 
Unpublished—Reid Moran 1961; E. L. Sleeper 1964. 
Pinus torreyana Parry MAP 55 
Torrey pine 
Pinus torreyana Parry ex Carr., Traite Gen. 
Conif. 326. 1855. 
Torrey pine has a very restricted range in southern 
California. On the mainland it is confined to the low coastal 
bluffs flanking the Soledad Valley north of San Diego.  Else- 
where it occurs only as a small grove on Santa Rosa Island. 
The mainland distribution shown on the map is based on 
Vegetation-Type mapping done in 1931; the present range 
of the species may be slightly less than that shown. 
Additional source: 
Unpublished—E. R. Blakley 1961 (Santa Rosa Island). 
