Unpublished—W. D. Billings 1961 (Nev.); T. C. Bray- 
shaw and A. E. Porsild 1964 (B.C.); W. P. Cottam 1964 
(Utah); J. D. Curtis 1964 (Utah) ; J. F. Franklin 1964 (Oreg.); 
J. R. Haller 1964 (Calif.); F. D. Johnson 1964 (Idaho); F. 
Kramer to N. T. Mirov (n.d.) (Nev.); O. V. Matthews 1964 
(Ores) eRe AY Read? 1964; R> Schmidt 1964 ((B:G.); W. 1: 
Stein 1964; R. F. Wagle 1962 (Ariz.); O. Zarzosa L. 1964 
(Dgo.). 
Pinus washoensis Mason & Stockwell MAP 48 
Washoe pine 
Pinus washoensis Mason & Stockwell, Madrono 
8: 62. 1945. 
Not distinguished until 1945, Washoe pine is known only 
from limited areas in three mountain ranges at the western 
edge of the Great Basin in Nevada and California. It oc- 
cupies a few square miles on the east slopes of Mt. Rose, Nev., 
and has recently been discovered in the southern Warner 
Mountains of northeastern California (Haller 1961) and on 
the Bald Mountain Range (Calif.), about 20 miles northwest 
of Mt. Rose (Critchfield and Allenbaugh 1965). In all three 
localities it grows at higher elevations than Pinus ponderosa, 
its closest relative. According to Haller, it may also be oc- 
casional in a variable population of P. ponderosa and P. 
jeffreyi near Hobart Mills, Calif., about 14 miles west of 
Mt. Rose. 
Sources: 
Published—Critchfield and Allenbaugh 1965; Haller 1961; 
Mason and Stockwell 1945. 
Pinus jeffreyi Grev. & Balf. MAP 48 
Jeffrey pine 
Pinus jeffreyi Grev. & Balf. in A. Murr., Bot. 
Exped. Oreg. [ Rpt. No. 8] 2, t. 1853. 
Primarily a California species, Jeffrey pine extends into 
southwestern Oregon, western Nevada, and northern Baja 
California. It usually grows at higher elevations than the 
closely related Pinus ponderosa, but the two overlap broadly 
and hybridize in nature (Haller 1962). Our map is based 
on a distribution map prepared by D. F. Roy and modified 
from the sources of information listed below. 
Additional sources: 
Published—Haller 1962. 
Thesis—Donald V. Hemphill 1952. The vertebrate 
fauna of the boreal areas of the southern Yolla Bolly Moun- 
MMe California. Ph.D. thesis, Oregon State College, 340 pp., 
illus. 
Unpublished—B. K. Ford 1963 (Calif.) ; J. R. Griffin 1964 
(Calif.); D. V. Hemphill 1964 (Calif.); J. L. Jenkinson 1963 
(Calif., Oreg.); Reid Moran 1964 (Baja Calif.). 
Pinus engelmannii Carr. MAP 49 
Apache pine 
Pinus engelmannii Carr., Rev. Hort., Sér. 4, 
3: 227. 1854; “engelmanni.” 
Apache pine extends from southeastern Arizona and ex- 
treme southwestern New Mexico through the Sierra Madre 
Occidental of Mexico to Zacatecas and Aguascalientes. 
This species has also been known as Pinus latifolia Sarg., 
P. apacheca Lemm., and P. macrophylla Engelm. It usually 
grows at lower elevations than the related yellow pines of this 
region (P. ponderosa, P. durangensis, P. cooper). 
Additional sources: 
Published—Gentry 1946 (Sin.); Guzman and Vela 1960 
(Zac., Ags.) ; J. 1. Marshall 1957 (Son., Chih.). 
Thesis—James H. Maysilles 1959. Floral relationships of 
the pine forests of western Durango, Mexico. Ph.D. thesis, 
Univ. Michigan, 165 pp., illus. 
Unpublished—O, Zarzosa L. 1964 (Dgo.); C. E. 
1964 (Dgo.). 
Blanco 
Pinus durangensis Martinez MAP 50 
Durango pine 
Pinus durangensis Martinez, Mex. Inst. Biol. An. 
ligre24, fio. 1-4. 1942 
This species is confined to Mexico, ranging from eastern 
Sonora to southern Durango, with outliers in Michoacan and 
Nuevo Leon. It is an important constituent of the yellow 
pine forests in the Sierra Madre Occidental. 
Additional sources: 
Published—J. T. Marshall 1957 
(Chih.) ; Zobel and Cech 1957 (N.L.). 
Thesis—James H. Maysilles 1959. Floral relationships 
of the pine forests of western Durango, Mexico. Ph.D. thesis, 
Univ. Michigan, 165 pp., illus. 
Unpublished—Mexico, Instituto Nacional de Investiga- 
ciones Forestales 1964 (Ags.); N. Sanchez Mejorada to L. W. 
Bryan 1963 (Mich.) ; O. Zarzosa L. 1964 (Dgo.). 
Wicht 1949 
5on.); 
Pinus cooperi C. E. Blanco MAP 51 
Cooper pine 
Pinus cooperi C. E. Blanco, Mex. Inst. Biol. An. 
202,185; -fig; 1_ 1949: 
This species grows at moderate elevations in the exten- 
sive pine forests of southern Chihuahua and western Durango. 
It is associated with several other species in the western yel- 
low pine group, including Pinus ponderosa (P. arizonica 
Engelm.) , P. engelmannii, and P. durangensis. P. coopert was 
originally named P. lutea C. E. Blanco (not Walt.) . 
Additional sources: 
Unpublished—C, E. Blanco 1964 (Dgo.); O. Zarzosa L. 
1964 (Dgo.); E. Hernandez X. and J. Vasquez Soto 1964 
(Chih.). 
Pinus montezumae Lamb. MAP 49 
Montezuma pine 
Pinus montezumae Lamb., Descr. Genus Pinus. Ed. 
3m(8 2) ls 59, tei22: 1832. 
This variable species is a major constituent of the pine 
forests of central Mexico, ranging from Jalisco to Veracruz 
and south to Oaxaca, Chiapas, and Guatemala. It is also 
present in the Sierra Madre Oriental, and its northernmost 
known occurrence is in an outlier of this range, the Sierra 
Picachos in Nuevo Leon (Zobel and Cech 1957). In various 
parts of its range it is associated with many of the other Mex- 
ican yellow pines, and it may intergrade with some of them 
(Loock 1950, p. 180) . 
Additional sources: 
Published—Aguilar 1961 (Guatemala); Goldman 1951; 
Hernandez X. et al. 1951 (Tamps.); Leavenworth 1946 
(Mich.) ; Loock 1950 (Mich.); Schwerdtfeger 1953 (Guate- 
mala) ; Zobel and Cech 1957 (N.L.). 
Unpublished—B. Hallberg 1964 (Oax.); E. Larsen 1962; 
F. Medellin-Leal 1964 (S.L.P.); N. Sanchez Mejorada to L. 
W. Bryan 1963 (Mich.). 
Pinus hartwegii Lindl. MAP 51 
Hartweg pine 
Pinus hartwegit Lindl., Bot. Reg. v. 25, Misc. 62. 1839. 
Pinus hartwegtt, including the doubtfully distinct P. 
rudis Endl., has a markedly discontinuous distribution at high 
elevations from Chihuahua and Nuevo Leén south to Guate- 
mala and El Salvador. It often reaches timberline, but at 
lower elevations it overlaps and may intergrade with P 
zumae (Standley and Steyermark 1958) . 
Additional sources: 
Published—Aguilar 1961 (Guatemala); Hinds and Lar 
sen 1961 (Mex.); Leavenworth 1946 (Mich Loock 1950 
